The native-stop safety atlas

A rescue that keeps going can create a second problem.

Readthrough should cross a premature stop—not a healthy protein's normal stop. This atlas measures that unwanted continuation for G418.

18,887normal stops represented by the model
1,700stops with prediction and measurement
1 drug · 1 cell lineempirical atlas coverage
01

Measurement coverage

Most stops could be predicted. Few could be measured.

The model scores one MANE reference transcript per gene. Direct comparison also needs enough ribosome reads around its normal stop.

Only 1,700 of 18,887 scored normal stops have a matched measurement

Prediction coverage versus prediction-and-measurement coverage, on the same denominator

measured
One normal stop codon per MANE transcriptGREY = NO MATCHED MEASUREMENTprediction + measurement: 1,700 canonical stops (9.0%)MATCHED1,7009.0%prediction only: 17,187 canonical stops (91.0%)NO MATCHED MEASUREMENT17,18791.0%CAN COMPARE MODEL WITH DATAPREDICTION EXISTS · EMPIRICAL CHECK DOES NOT

BLUE CAN TEST THE MODEL · Grey is missing empirical evidence, not evidence of zero native-stop readthrough.

Details & limits
The matched subset is 9.0% of the scored atlas. Stops outside it were not expressed or sequenced deeply enough for this comparison in HEK293T; they are not assumed to behave like the matched set.
9.0%directly checked

All later results use these 1,700 stops, in one cell line.

02

Prediction versus measurement

The model sees a signal. It cannot replace the assay.

Each dot is one normal stop. Right means higher prediction; up means more measured continuation after G418.

Predicted and measured native-stop burden have weak-to-moderate rank agreement

1,700 matched genes · G418 versus control · HEK293T

measured
0.0%1.0%2.0%3.0%4.0%−2 pp0 pp+5 pp+10 pp+15 ppARF5: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothFKBP4: predicted G418 readthrough 1.1%; measured change after the stop +1.37 percentage points; High in model onlyRPAP3: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyNDUFAB1: predicted G418 readthrough 0.3%; measured change after the stop −0.24 percentage points; Lower in bothPPP5C: predicted G418 readthrough 0.5%; measured change after the stop +0.13 percentage points; Lower in bothCOX15: predicted G418 readthrough 1.1%; measured change after the stop +3.70 percentage points; High in model onlyVDAC3: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothAKAP11: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothGABARAPL2: predicted G418 readthrough 0.9%; measured change after the stop +2.64 percentage points; High in model onlyMRI1: predicted G418 readthrough 0.5%; measured change after the stop +0.48 percentage points; Lower in bothBOD1L1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothPSMA4: predicted G418 readthrough 0.5%; measured change after the stop +3.18 percentage points; Lower in bothGRN: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothVAMP3: predicted G418 readthrough 0.6%; measured change after the stop +8.09 percentage points; High in measurement onlyMRPS35: predicted G418 readthrough 0.5%; measured change after the stop +0.14 percentage points; Lower in bothPSMC4: predicted G418 readthrough 2.0%; measured change after the stop +2.90 percentage points; High in model onlyTRIP13: predicted G418 readthrough 0.9%; measured change after the stop +3.32 percentage points; High in model onlyHSD17B10: predicted G418 readthrough 0.7%; measured change after the stop +1.05 percentage points; Lower in bothISOC1: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothLAPTM4A: predicted G418 readthrough 0.8%; measured change after the stop +1.03 percentage points; Lower in bothNDUFB4: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothTMEM131: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothBLTP3A: predicted G418 readthrough 1.3%; measured change after the stop +2.82 percentage points; High in model onlyRPS5: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyRBM22: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; High in model onlyLAMP2: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothDLD: predicted G418 readthrough 1.0%; measured change after the stop +1.71 percentage points; High in model onlyCDC6: predicted G418 readthrough 0.3%; measured change after the stop +0.81 percentage points; Lower in bothVCL: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothPSMD8: predicted G418 readthrough 0.6%; measured change after the stop +1.13 percentage points; Lower in bothATP5F1D: predicted G418 readthrough 0.3%; measured change after the stop +1.07 percentage points; Lower in bothNDUFB7: predicted G418 readthrough 0.5%; measured change after the stop +2.39 percentage points; Lower in bothTIMM13: predicted G418 readthrough 1.7%; measured change after the stop +2.60 percentage points; High in model onlyMIF: predicted G418 readthrough 0.4%; measured change after the stop +0.23 percentage points; Lower in bothDDTL: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothSF3A1: predicted G418 readthrough 0.3%; measured change after the stop +1.05 percentage points; Lower in bothMAPK1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothTOMM22: predicted G418 readthrough 0.4%; measured change after the stop +0.29 percentage points; Lower in bothRTCB: predicted G418 readthrough 0.6%; measured change after the stop +2.65 percentage points; Lower in bothHMOX1: predicted G418 readthrough 0.7%; measured change after the stop +1.32 percentage points; Lower in bothMCM5: predicted G418 readthrough 0.6%; measured change after the stop +1.91 percentage points; Lower in bothTTLL12: predicted G418 readthrough 0.3%; measured change after the stop +1.00 percentage points; Lower in bothRPL3: predicted G418 readthrough 0.6%; measured change after the stop +0.26 percentage points; Lower in bothMYH9: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothEIF3D: predicted G418 readthrough 0.3%; measured change after the stop +1.27 percentage points; Lower in bothST13: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothRBX1: predicted G418 readthrough 0.5%; measured change after the stop +5.19 percentage points; High in measurement onlyPHF5A: predicted G418 readthrough 1.1%; measured change after the stop +17.99 percentage points; High in model + measurementHSP90AA1: predicted G418 readthrough 0.5%; measured change after the stop +0.19 percentage points; Lower in bothNID2: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothSUPT16H: predicted G418 readthrough 0.3%; measured change after the stop +0.75 percentage points; Lower in bothPYGL: predicted G418 readthrough 1.2%; measured change after the stop +2.00 percentage points; High in model onlyCNIH1: predicted G418 readthrough 0.6%; measured change after the stop −0.82 percentage points; Lower in bothPSMA3: predicted G418 readthrough 0.6%; measured change after the stop +2.38 percentage points; Lower in bothAHSA1: predicted G418 readthrough 0.9%; measured change after the stop +0.00 percentage points; High in model onlyPABPN1: predicted G418 readthrough 0.4%; measured change after the stop −0.36 percentage points; Lower in bothPNN: predicted G418 readthrough 0.4%; measured change after the stop +0.63 percentage points; Lower in bothCHMP4B: predicted G418 readthrough 0.3%; measured change after the stop +0.52 percentage points; Lower in bothAHCY: predicted G418 readthrough 0.7%; measured change after the stop +1.11 percentage points; Lower in bothPSMD10: predicted G418 readthrough 0.3%; measured change after the stop +1.06 percentage points; Lower in bothPGRMC1: predicted G418 readthrough 0.5%; measured change after the stop +0.55 percentage points; Lower in bothABCD1: predicted G418 readthrough 2.2%; measured change after the stop +1.83 percentage points; High in model onlyRP2: predicted G418 readthrough 0.4%; measured change after the stop +2.59 percentage points; Lower in bothHTATSF1: predicted G418 readthrough 0.4%; measured change after the stop +0.61 percentage points; Lower in bothPOLR2C: predicted G418 readthrough 0.8%; measured change after the stop +1.21 percentage points; Lower in bothPSMD7: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothEIF3E: predicted G418 readthrough 0.5%; measured change after the stop +1.24 percentage points; Lower in bothNEFM: predicted G418 readthrough 0.4%; measured change after the stop −0.29 percentage points; Lower in bothHNRNPL: predicted G418 readthrough 0.7%; measured change after the stop +0.16 percentage points; Lower in bothRNASEH2A: predicted G418 readthrough 1.4%; measured change after the stop +1.47 percentage points; High in model onlySF3A2: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothFBL: predicted G418 readthrough 0.6%; measured change after the stop +1.30 percentage points; Lower in bothCDC37: predicted G418 readthrough 1.1%; measured change after the stop +0.94 percentage points; High in model onlyLENG1: predicted G418 readthrough 1.7%; measured change after the stop +0.85 percentage points; High in model onlyRPL18A: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothPOLR1F: predicted G418 readthrough 0.7%; measured change after the stop +0.09 percentage points; Lower in bothHOXA5: predicted G418 readthrough 0.7%; measured change after the stop +2.21 percentage points; Lower in bothAIMP2: predicted G418 readthrough 0.5%; measured change after the stop +0.20 percentage points; Lower in bothRPA3: predicted G418 readthrough 1.5%; measured change after the stop +0.00 percentage points; High in model onlyPSMA2: predicted G418 readthrough 0.9%; measured change after the stop +0.57 percentage points; High in model onlySEC61B: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothPPIF: predicted G418 readthrough 0.6%; measured change after the stop +4.39 percentage points; Lower in bothDHX33: predicted G418 readthrough 0.6%; measured change after the stop +1.04 percentage points; Lower in bothNUFIP2: predicted G418 readthrough 0.8%; measured change after the stop +0.75 percentage points; Lower in bothRPL19: predicted G418 readthrough 0.4%; measured change after the stop +0.16 percentage points; Lower in bothPFN1: predicted G418 readthrough 1.6%; measured change after the stop +0.00 percentage points; High in model onlySLC25A11: predicted G418 readthrough 0.6%; measured change after the stop +1.16 percentage points; Lower in bothC1QBP: predicted G418 readthrough 0.3%; measured change after the stop +0.94 percentage points; Lower in bothCPD: predicted G418 readthrough 1.3%; measured change after the stop +1.02 percentage points; High in model onlyCCDC47: predicted G418 readthrough 0.3%; measured change after the stop +0.93 percentage points; Lower in bothDDX5: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothMRPL27: predicted G418 readthrough 1.6%; measured change after the stop +1.59 percentage points; High in model onlyLRRC59: predicted G418 readthrough 0.4%; measured change after the stop +0.81 percentage points; Lower in bothTMEM97: predicted G418 readthrough 0.7%; measured change after the stop +5.37 percentage points; High in measurement onlyLAP3: predicted G418 readthrough 0.5%; measured change after the stop +0.85 percentage points; Lower in bothCCDC86: predicted G418 readthrough 0.5%; measured change after the stop +1.65 percentage points; Lower in bothPRPF19: predicted G418 readthrough 0.4%; measured change after the stop +1.16 percentage points; Lower in bothTMEM109: predicted G418 readthrough 0.5%; measured change after the stop +2.10 percentage points; Lower in bothC11orf58: predicted G418 readthrough 0.5%; measured change after the stop −0.70 percentage points; Lower in bothPOLR3B: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothARPC3: predicted G418 readthrough 0.5%; measured change after the stop +2.97 percentage points; Lower in bothGPN3: predicted G418 readthrough 0.6%; measured change after the stop −0.69 percentage points; Lower in bothKRR1: predicted G418 readthrough 0.6%; measured change after the stop +1.61 percentage points; Lower in bothMRPL51: predicted G418 readthrough 0.4%; measured change after the stop +1.94 percentage points; Lower in bothGAPDH: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothCOX6A1: predicted G418 readthrough 0.3%; measured change after the stop +2.01 percentage points; Lower in bothSRSF9: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothASF1A: predicted G418 readthrough 1.8%; measured change after the stop +5.19 percentage points; High in model + measurementFBXO5: predicted G418 readthrough 0.9%; measured change after the stop +1.24 percentage points; High in model onlyBYSL: predicted G418 readthrough 0.4%; measured change after the stop +1.20 percentage points; Lower in bothDNPH1: predicted G418 readthrough 1.4%; measured change after the stop +1.47 percentage points; High in model onlyMTREX: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothNUP155: predicted G418 readthrough 0.8%; measured change after the stop +0.51 percentage points; High in model onlyC5orf15: predicted G418 readthrough 0.5%; measured change after the stop −0.81 percentage points; Lower in bothTRIM23: predicted G418 readthrough 1.5%; measured change after the stop +0.79 percentage points; High in model onlyRARS1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothHES1: predicted G418 readthrough 0.6%; measured change after the stop +0.74 percentage points; Lower in bothUMPS: predicted G418 readthrough 0.8%; measured change after the stop +0.83 percentage points; High in model onlyDDX1: predicted G418 readthrough 1.5%; measured change after the stop +0.65 percentage points; High in model onlyTMSB10: predicted G418 readthrough 0.4%; measured change after the stop +5.98 percentage points; High in measurement onlyMSH2: predicted G418 readthrough 0.6%; measured change after the stop +0.98 percentage points; Lower in bothSF3B6: predicted G418 readthrough 0.4%; measured change after the stop +0.38 percentage points; Lower in bothHSPE1: predicted G418 readthrough 1.1%; measured change after the stop +2.28 percentage points; High in model onlyCEBPZ: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothDLX2: predicted G418 readthrough 1.0%; measured change after the stop +0.40 percentage points; High in model onlyPPP3R1: predicted G418 readthrough 1.2%; measured change after the stop +0.98 percentage points; High in model onlyMSH6: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothENO1: predicted G418 readthrough 0.2%; measured change after the stop +0.05 percentage points; Lower in bothRPL22: predicted G418 readthrough 0.4%; measured change after the stop +2.98 percentage points; Lower in bothWDR77: predicted G418 readthrough 0.3%; measured change after the stop +0.65 percentage points; Lower in bothAKR7A2: predicted G418 readthrough 0.5%; measured change after the stop +1.55 percentage points; Lower in bothEBNA1BP2: predicted G418 readthrough 0.4%; measured change after the stop +0.10 percentage points; Lower in bothCTSD: predicted G418 readthrough 0.6%; measured change after the stop +0.78 percentage points; Lower in bothATIC: predicted G418 readthrough 1.0%; measured change after the stop +0.18 percentage points; High in model onlyMED28: predicted G418 readthrough 0.4%; measured change after the stop +3.60 percentage points; Lower in bothYWHAQ: predicted G418 readthrough 0.3%; measured change after the stop +2.71 percentage points; Lower in bothCPSF3: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothIRF2BPL: predicted G418 readthrough 0.7%; measured change after the stop +1.02 percentage points; Lower in bothFAM98A: predicted G418 readthrough 1.5%; measured change after the stop +2.75 percentage points; High in model onlyYIPF4: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothMSX2: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothTARDBP: predicted G418 readthrough 0.3%; measured change after the stop +1.42 percentage points; Lower in bothCOIL: predicted G418 readthrough 1.1%; measured change after the stop +0.00 percentage points; High in model onlyNAA50: predicted G418 readthrough 0.6%; measured change after the stop +0.53 percentage points; Lower in bothMRPS2: predicted G418 readthrough 1.0%; measured change after the stop +6.60 percentage points; High in model + measurementZC3HAV1: predicted G418 readthrough 0.9%; measured change after the stop +0.00 percentage points; High in model onlyTRIR: predicted G418 readthrough 2.0%; measured change after the stop +11.36 percentage points; High in model + measurementLRP1: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothSEC61A1: predicted G418 readthrough 1.0%; measured change after the stop +1.95 percentage points; High in model onlyATP5F1E: predicted G418 readthrough 0.4%; measured change after the stop −0.27 percentage points; Lower in bothSRSF6: predicted G418 readthrough 0.7%; measured change after the stop +1.60 percentage points; Lower in bothMOCS3: predicted G418 readthrough 0.6%; measured change after the stop +2.63 percentage points; Lower in bothMPHOSPH10: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothSNRPC: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothH1-3: predicted G418 readthrough 0.8%; measured change after the stop +0.62 percentage points; Lower in bothMEA1: predicted G418 readthrough 1.7%; measured change after the stop +1.31 percentage points; High in model onlySOX4: predicted G418 readthrough 0.7%; measured change after the stop +0.77 percentage points; Lower in bothGOT2: predicted G418 readthrough 0.3%; measured change after the stop +0.74 percentage points; Lower in bothRPS29: predicted G418 readthrough 0.5%; measured change after the stop +2.78 percentage points; Lower in bothMRPS7: predicted G418 readthrough 0.3%; measured change after the stop +3.01 percentage points; Lower in bothCLPP: predicted G418 readthrough 0.7%; measured change after the stop +2.25 percentage points; Lower in bothDSTN: predicted G418 readthrough 0.3%; measured change after the stop +1.21 percentage points; Lower in bothSNRPB2: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothCAPNS1: predicted G418 readthrough 1.1%; measured change after the stop +3.01 percentage points; High in model onlyUBA2: predicted G418 readthrough 1.0%; measured change after the stop +3.76 percentage points; High in model onlyARL2: predicted G418 readthrough 1.0%; measured change after the stop +3.36 percentage points; High in model onlyTRAP1: predicted G418 readthrough 2.2%; measured change after the stop +0.00 percentage points; High in model onlyHSPA2: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothCANX: predicted G418 readthrough 0.6%; measured change after the stop +0.11 percentage points; Lower in bothCOX7C: predicted G418 readthrough 0.2%; measured change after the stop −0.47 percentage points; Lower in bothJPT2: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothEMC6: predicted G418 readthrough 0.8%; measured change after the stop +11.82 percentage points; High in measurement onlySDF2L1: predicted G418 readthrough 1.1%; measured change after the stop −0.07 percentage points; High in model onlyYWHAH: predicted G418 readthrough 0.5%; measured change after the stop +9.39 percentage points; High in measurement onlyATP6V1F: predicted G418 readthrough 0.5%; measured change after the stop +1.15 percentage points; Lower in bothHOXD11: predicted G418 readthrough 1.0%; measured change after the stop +6.95 percentage points; High in model + measurementTXNDC17: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothFXR2: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothDAD1: predicted G418 readthrough 0.6%; measured change after the stop +1.37 percentage points; Lower in bothLMAN1: predicted G418 readthrough 3.5%; measured change after the stop +3.03 percentage points; High in model onlyMEGF8: predicted G418 readthrough 3.0%; measured change after the stop +1.24 percentage points; High in model onlyRPS16: predicted G418 readthrough 0.3%; measured change after the stop +0.33 percentage points; Lower in bothNCAPG: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlySCML2: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothNDUFA2: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlySAFB2: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothCOLGALT1: predicted G418 readthrough 0.9%; measured change after the stop +1.80 percentage points; High in model onlyMLLT1: predicted G418 readthrough 1.9%; measured change after the stop +3.20 percentage points; High in model onlyACTN4: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothJUND: predicted G418 readthrough 0.7%; measured change after the stop +2.38 percentage points; Lower in bothTAF4: predicted G418 readthrough 1.9%; measured change after the stop +0.66 percentage points; High in model onlyLAMA5: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothUBE2M: predicted G418 readthrough 0.4%; measured change after the stop +4.23 percentage points; Lower in bothTRIM28: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothEIF2S3: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothZC3H4: predicted G418 readthrough 0.3%; measured change after the stop +1.08 percentage points; Lower in bothEIF3G: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothGRWD1: predicted G418 readthrough 0.7%; measured change after the stop +0.40 percentage points; Lower in bothNOL11: predicted G418 readthrough 2.0%; measured change after the stop +0.27 percentage points; High in model onlyCOX4I1: predicted G418 readthrough 0.6%; measured change after the stop +3.31 percentage points; Lower in bothGINS2: predicted G418 readthrough 0.6%; measured change after the stop +0.86 percentage points; Lower in bothVPS25: predicted G418 readthrough 1.4%; measured change after the stop +1.20 percentage points; High in model onlyFUS: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothNUP210: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothMYBBP1A: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothRPA1: predicted G418 readthrough 0.6%; measured change after the stop +1.64 percentage points; Lower in bothGRSF1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothUTP3: predicted G418 readthrough 0.7%; measured change after the stop +0.34 percentage points; Lower in bothH3-3B: predicted G418 readthrough 0.5%; measured change after the stop +0.83 percentage points; Lower in bothNIP7: predicted G418 readthrough 0.3%; measured change after the stop +2.42 percentage points; Lower in bothEIF2S1: predicted G418 readthrough 0.6%; measured change after the stop +0.16 percentage points; Lower in bothKGD4: predicted G418 readthrough 0.6%; measured change after the stop +2.91 percentage points; Lower in bothCCNB1: predicted G418 readthrough 0.3%; measured change after the stop +0.46 percentage points; Lower in bothNOTCH2: predicted G418 readthrough 0.8%; measured change after the stop +0.30 percentage points; Lower in bothSLC38A2: predicted G418 readthrough 0.5%; measured change after the stop +1.51 percentage points; Lower in bothRTL8C: predicted G418 readthrough 0.7%; measured change after the stop +5.59 percentage points; High in measurement onlyPRPF38A: predicted G418 readthrough 0.3%; measured change after the stop +0.70 percentage points; Lower in bothCDK4: predicted G418 readthrough 0.4%; measured change after the stop +2.44 percentage points; Lower in bothESPL1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothCCT7: predicted G418 readthrough 0.7%; measured change after the stop +0.45 percentage points; Lower in bothURB2: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothLAMC1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothMRPL44: predicted G418 readthrough 1.1%; measured change after the stop +1.42 percentage points; High in model onlyCOX5B: predicted G418 readthrough 0.5%; measured change after the stop +2.17 percentage points; Lower in bothCKAP2: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothKDELR2: predicted G418 readthrough 0.4%; measured change after the stop +1.41 percentage points; Lower in bothIREB2: predicted G418 readthrough 0.6%; measured change after the stop +0.91 percentage points; Lower in bothSRSF1: predicted G418 readthrough 0.2%; measured change after the stop +0.06 percentage points; Lower in bothPSMB7: predicted G418 readthrough 1.0%; measured change after the stop +1.86 percentage points; High in model onlyRANBP6: predicted G418 readthrough 1.0%; measured change after the stop +0.77 percentage points; High in model onlyTUBB2B: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothMRPS18B: predicted G418 readthrough 0.4%; measured change after the stop +1.46 percentage points; Lower in bothMRPL15: predicted G418 readthrough 0.7%; measured change after the stop −0.23 percentage points; Lower in bothRPLP1: predicted G418 readthrough 0.4%; measured change after the stop +0.02 percentage points; Lower in bothACTR2: predicted G418 readthrough 0.3%; measured change after the stop +0.70 percentage points; Lower in bothKIF11: predicted G418 readthrough 0.5%; measured change after the stop +0.42 percentage points; Lower in bothDBR1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothTOPBP1: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothDNAJC13: predicted G418 readthrough 0.8%; measured change after the stop −0.44 percentage points; Lower in bothSSB: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothWBP11: predicted G418 readthrough 2.0%; measured change after the stop +5.83 percentage points; High in model + measurementTRMT5: predicted G418 readthrough 0.3%; measured change after the stop +0.12 percentage points; Lower in bothPEBP1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothZNF24: predicted G418 readthrough 0.3%; measured change after the stop −0.59 percentage points; Lower in bothLMNB1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothNUP133: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothPSMA6: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothUSP22: predicted G418 readthrough 0.5%; measured change after the stop +2.11 percentage points; Lower in bothMIB1: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyDSG2: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothRNF138: predicted G418 readthrough 0.6%; measured change after the stop +0.60 percentage points; Lower in bothNDC80: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothUSP14: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothHERC2: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothSLC7A5: predicted G418 readthrough 0.7%; measured change after the stop +0.38 percentage points; Lower in bothCYBA: predicted G418 readthrough 1.1%; measured change after the stop +0.00 percentage points; High in model onlyUTP20: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothKPNA3: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothRTRAF: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothBLMH: predicted G418 readthrough 1.1%; measured change after the stop +0.00 percentage points; High in model onlyERP29: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothHACD3: predicted G418 readthrough 0.4%; measured change after the stop +0.61 percentage points; Lower in bothRAB11A: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothFAF2: predicted G418 readthrough 2.1%; measured change after the stop +0.00 percentage points; High in model onlyRBM25: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothATP5F1B: predicted G418 readthrough 0.5%; measured change after the stop +0.72 percentage points; Lower in bothPTGES3: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothPHF3: predicted G418 readthrough 0.3%; measured change after the stop +0.88 percentage points; Lower in bothEIF4B: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothCOPZ1: predicted G418 readthrough 0.5%; measured change after the stop +0.89 percentage points; Lower in bothNUP153: predicted G418 readthrough 0.3%; measured change after the stop +0.75 percentage points; Lower in bothPSMB1: predicted G418 readthrough 1.4%; measured change after the stop +1.50 percentage points; High in model onlyYY1: predicted G418 readthrough 0.6%; measured change after the stop +0.96 percentage points; Lower in bothWLS: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothCREBBP: predicted G418 readthrough 0.9%; measured change after the stop +1.28 percentage points; High in model onlyCOTL1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothSLC12A2: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothFBN2: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; High in model onlyCFAP20: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothSRCAP: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothSETD1A: predicted G418 readthrough 0.6%; measured change after the stop +0.05 percentage points; Lower in bothZNF629: predicted G418 readthrough 0.8%; measured change after the stop +0.70 percentage points; Lower in bothUPF1: predicted G418 readthrough 0.4%; measured change after the stop +0.78 percentage points; Lower in bothMYDGF: predicted G418 readthrough 1.9%; measured change after the stop +9.19 percentage points; High in model + measurementPTPRS: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothCSE1L: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothCCDC6: predicted G418 readthrough 0.4%; measured change after the stop +0.95 percentage points; Lower in bothCABIN1: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothCAPZA1: predicted G418 readthrough 0.5%; measured change after the stop +1.20 percentage points; Lower in bothCDC45: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothDDX18: predicted G418 readthrough 0.7%; measured change after the stop −0.39 percentage points; Lower in bothEP300: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothAP2S1: predicted G418 readthrough 0.4%; measured change after the stop −0.56 percentage points; Lower in bothPOLR1B: predicted G418 readthrough 0.4%; measured change after the stop +0.42 percentage points; Lower in bothASF1B: predicted G418 readthrough 0.5%; measured change after the stop +5.35 percentage points; High in measurement onlyWDR11: predicted G418 readthrough 1.6%; measured change after the stop +0.00 percentage points; High in model onlyVPS26A: predicted G418 readthrough 0.7%; measured change after the stop +0.95 percentage points; Lower in bothCD81: predicted G418 readthrough 0.7%; measured change after the stop +2.21 percentage points; Lower in bothPNO1: predicted G418 readthrough 1.5%; measured change after the stop +5.38 percentage points; High in model + measurementTAF1B: predicted G418 readthrough 1.1%; measured change after the stop +3.04 percentage points; High in model onlySPCS2: predicted G418 readthrough 0.7%; measured change after the stop +0.43 percentage points; Lower in bothSNRNP40: predicted G418 readthrough 0.4%; measured change after the stop +1.78 percentage points; Lower in bothDNAJC8: predicted G418 readthrough 1.4%; measured change after the stop +2.50 percentage points; High in model onlySRBD1: predicted G418 readthrough 1.1%; measured change after the stop +0.61 percentage points; High in model onlyPOLR1A: predicted G418 readthrough 0.6%; measured change after the stop +0.27 percentage points; Lower in bothSC5D: predicted G418 readthrough 0.4%; measured change after the stop +2.16 percentage points; Lower in bothARCN1: predicted G418 readthrough 0.5%; measured change after the stop +0.78 percentage points; Lower in bothHAT1: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothMCM6: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyDARS1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothNID1: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothPPAT: predicted G418 readthrough 0.9%; measured change after the stop +0.84 percentage points; High in model onlyRPL31: predicted G418 readthrough 0.2%; measured change after the stop +0.49 percentage points; Lower in bothNOP58: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyYWHAE: predicted G418 readthrough 0.6%; measured change after the stop +0.89 percentage points; Lower in bothUBE2S: predicted G418 readthrough 0.4%; measured change after the stop +0.18 percentage points; Lower in bothPSMD3: predicted G418 readthrough 0.5%; measured change after the stop +2.90 percentage points; Lower in bothCAD: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothSEPTIN11: predicted G418 readthrough 0.3%; measured change after the stop +1.50 percentage points; Lower in bothGRPEL1: predicted G418 readthrough 1.5%; measured change after the stop +3.47 percentage points; High in model onlyMRPL3: predicted G418 readthrough 1.0%; measured change after the stop +0.99 percentage points; High in model onlySSR3: predicted G418 readthrough 0.6%; measured change after the stop +2.61 percentage points; Lower in bothMCM2: predicted G418 readthrough 1.0%; measured change after the stop +3.51 percentage points; High in model onlyZFR: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothSUB1: predicted G418 readthrough 0.3%; measured change after the stop +2.65 percentage points; Lower in bothVCAN: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlySTC2: predicted G418 readthrough 0.8%; measured change after the stop +5.03 percentage points; High in measurement onlyRPL26L1: predicted G418 readthrough 0.4%; measured change after the stop +0.96 percentage points; Lower in bothCENPE: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothLSG1: predicted G418 readthrough 0.5%; measured change after the stop +1.84 percentage points; Lower in bothRBM27: predicted G418 readthrough 0.6%; measured change after the stop +0.76 percentage points; Lower in bothSPDL1: predicted G418 readthrough 0.2%; measured change after the stop +0.93 percentage points; Lower in bothSSBP1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothVDAC1: predicted G418 readthrough 0.5%; measured change after the stop +0.97 percentage points; Lower in bothAFF4: predicted G418 readthrough 1.1%; measured change after the stop +1.26 percentage points; High in model onlyXPO5: predicted G418 readthrough 0.7%; measured change after the stop +0.37 percentage points; Lower in bothEIF4H: predicted G418 readthrough 0.7%; measured change after the stop +2.60 percentage points; Lower in bothSQLE: predicted G418 readthrough 0.4%; measured change after the stop −0.26 percentage points; Lower in bothBTAF1: predicted G418 readthrough 0.7%; measured change after the stop +0.62 percentage points; Lower in bothTMEM115: predicted G418 readthrough 0.5%; measured change after the stop +0.60 percentage points; Lower in bothEIF2B2: predicted G418 readthrough 2.9%; measured change after the stop +1.29 percentage points; High in model onlyALG10: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothNDUFA9: predicted G418 readthrough 0.5%; measured change after the stop +0.78 percentage points; Lower in bothTMTC3: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothSNRPF: predicted G418 readthrough 1.1%; measured change after the stop +2.23 percentage points; High in model onlyTMBIM6: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothANKRD52: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothZIC5: predicted G418 readthrough 0.7%; measured change after the stop +0.80 percentage points; Lower in bothNOP9: predicted G418 readthrough 0.9%; measured change after the stop +0.63 percentage points; High in model onlySRP14: predicted G418 readthrough 0.3%; measured change after the stop +0.40 percentage points; Lower in bothTSPAN3: predicted G418 readthrough 0.6%; measured change after the stop +0.41 percentage points; Lower in bothTICRR: predicted G418 readthrough 1.2%; measured change after the stop +1.61 percentage points; High in model onlyUQCRC2: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothMAP1LC3B: predicted G418 readthrough 0.2%; measured change after the stop +0.44 percentage points; Lower in bothNDUFB10: predicted G418 readthrough 1.0%; measured change after the stop +0.37 percentage points; High in model onlyNOB1: predicted G418 readthrough 0.5%; measured change after the stop +0.59 percentage points; Lower in bothCLTC: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothCDH2: predicted G418 readthrough 1.1%; measured change after the stop +0.62 percentage points; High in model onlySLC39A6: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothARHGDIA: predicted G418 readthrough 0.9%; measured change after the stop +1.62 percentage points; High in model onlyFEM1A: predicted G418 readthrough 1.7%; measured change after the stop +1.62 percentage points; High in model onlySH3GL1: predicted G418 readthrough 1.3%; measured change after the stop +1.52 percentage points; High in model onlySOD1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothTIMM29: predicted G418 readthrough 0.6%; measured change after the stop +8.42 percentage points; High in measurement onlyPSMB6: predicted G418 readthrough 1.0%; measured change after the stop +1.26 percentage points; High in model onlyRPS11: predicted G418 readthrough 1.0%; measured change after the stop −0.12 percentage points; High in model onlyPGD: predicted G418 readthrough 1.5%; measured change after the stop +2.07 percentage points; High in model onlyPSMA5: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothCELSR2: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlySFT2D2: predicted G418 readthrough 0.4%; measured change after the stop +1.68 percentage points; Lower in bothSF3B4: predicted G418 readthrough 0.2%; measured change after the stop +0.17 percentage points; Lower in bothS100A11: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothPOGZ: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothSDE2: predicted G418 readthrough 0.8%; measured change after the stop +1.12 percentage points; Lower in bothARF1: predicted G418 readthrough 0.6%; measured change after the stop +3.43 percentage points; Lower in bothLBR: predicted G418 readthrough 0.4%; measured change after the stop −0.41 percentage points; Lower in bothPDIA6: predicted G418 readthrough 0.6%; measured change after the stop +0.99 percentage points; Lower in bothRHOB: predicted G418 readthrough 0.9%; measured change after the stop +9.10 percentage points; High in model + measurementCALM2: predicted G418 readthrough 0.5%; measured change after the stop +3.27 percentage points; Lower in bothRPS27A: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothSNRPG: predicted G418 readthrough 0.4%; measured change after the stop +0.06 percentage points; Lower in bothDYNC1LI1: predicted G418 readthrough 0.7%; measured change after the stop +2.07 percentage points; Lower in bothARL6IP5: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothLIMD1: predicted G418 readthrough 1.1%; measured change after the stop +0.77 percentage points; High in model onlyABHD10: predicted G418 readthrough 0.6%; measured change after the stop +0.16 percentage points; Lower in bothATP6V1A: predicted G418 readthrough 0.7%; measured change after the stop +0.63 percentage points; Lower in bothCCNA2: predicted G418 readthrough 0.7%; measured change after the stop +0.78 percentage points; Lower in bothNDUFS6: predicted G418 readthrough 0.4%; measured change after the stop +3.68 percentage points; Lower in bothMARCHF6: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothRPL37: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothNHP2: predicted G418 readthrough 0.9%; measured change after the stop +1.50 percentage points; High in model onlyZMAT2: predicted G418 readthrough 0.7%; measured change after the stop +0.66 percentage points; Lower in bothHIGD2A: predicted G418 readthrough 0.3%; measured change after the stop +0.74 percentage points; Lower in bothABT1: predicted G418 readthrough 0.8%; measured change after the stop +0.93 percentage points; Lower in bothPM20D2: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothCCT6A: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothZC3HAV1L: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothDOCK11: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothNDUFB9: predicted G418 readthrough 0.4%; measured change after the stop +1.13 percentage points; Lower in bothFAM120A: predicted G418 readthrough 0.2%; measured change after the stop +0.33 percentage points; Lower in bothPROSER2: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothLIN7C: predicted G418 readthrough 0.5%; measured change after the stop +5.19 percentage points; High in measurement onlySSRP1: predicted G418 readthrough 0.3%; measured change after the stop +1.16 percentage points; Lower in bothMTA2: predicted G418 readthrough 0.7%; measured change after the stop +3.02 percentage points; Lower in bothVPS51: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothCCT5: predicted G418 readthrough 0.6%; measured change after the stop +0.29 percentage points; Lower in bothDENR: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothNOCT: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothNDUFC2: predicted G418 readthrough 0.7%; measured change after the stop +0.94 percentage points; Lower in bothMZT2B: predicted G418 readthrough 0.5%; measured change after the stop −0.13 percentage points; Lower in bothOBI1: predicted G418 readthrough 3.7%; measured change after the stop −0.34 percentage points; High in model onlyMARS2: predicted G418 readthrough 0.4%; measured change after the stop +1.17 percentage points; Lower in bothGJA1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothRIOK2: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothSMARCA5: predicted G418 readthrough 1.2%; measured change after the stop +1.09 percentage points; High in model onlyRANBP2: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothRMND5A: predicted G418 readthrough 0.8%; measured change after the stop +1.42 percentage points; Lower in bothDDAH1: predicted G418 readthrough 0.7%; measured change after the stop −0.56 percentage points; Lower in bothTOMM70: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothUCHL1: predicted G418 readthrough 0.6%; measured change after the stop +1.35 percentage points; Lower in bothOLA1: predicted G418 readthrough 0.3%; measured change after the stop +0.05 percentage points; Lower in bothATP5MC3: predicted G418 readthrough 0.7%; measured change after the stop +0.11 percentage points; Lower in bothADAMTS1: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothCA2: predicted G418 readthrough 0.3%; measured change after the stop +1.37 percentage points; Lower in bothHSPA13: predicted G418 readthrough 0.6%; measured change after the stop −0.75 percentage points; Lower in bothNIFK: predicted G418 readthrough 0.8%; measured change after the stop +0.83 percentage points; Lower in bothGEMIN5: predicted G418 readthrough 0.6%; measured change after the stop +0.92 percentage points; Lower in bothAKR1B1: predicted G418 readthrough 1.1%; measured change after the stop +0.56 percentage points; High in model onlyNUP205: predicted G418 readthrough 0.7%; measured change after the stop +0.34 percentage points; Lower in bothFZD7: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothATP1B3: predicted G418 readthrough 0.8%; measured change after the stop +0.58 percentage points; Lower in bothVBP1: predicted G418 readthrough 0.5%; measured change after the stop −0.34 percentage points; Lower in bothCCT8: predicted G418 readthrough 1.2%; measured change after the stop +1.00 percentage points; High in model onlyUQCRB: predicted G418 readthrough 0.3%; measured change after the stop +0.36 percentage points; Lower in bothRPL30: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothSASS6: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothNOMO1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothABCF2: predicted G418 readthrough 0.4%; measured change after the stop +0.22 percentage points; Lower in bothCCNB2: predicted G418 readthrough 0.5%; measured change after the stop +0.77 percentage points; Lower in bothMRPL17: predicted G418 readthrough 0.6%; measured change after the stop +6.69 percentage points; High in measurement onlyEIF5B: predicted G418 readthrough 0.3%; measured change after the stop +0.15 percentage points; Lower in bothMIS18A: predicted G418 readthrough 1.3%; measured change after the stop +0.00 percentage points; High in model onlyKPNB1: predicted G418 readthrough 1.4%; measured change after the stop +0.45 percentage points; High in model onlyNCOA5: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothATP5PO: predicted G418 readthrough 0.3%; measured change after the stop +1.16 percentage points; Lower in bothIGF2BP1: predicted G418 readthrough 1.7%; measured change after the stop +1.29 percentage points; High in model onlyCBR1: predicted G418 readthrough 0.5%; measured change after the stop −0.04 percentage points; Lower in bothPSMB4: predicted G418 readthrough 0.9%; measured change after the stop +1.07 percentage points; High in model onlyHK2: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothATP6V0D1: predicted G418 readthrough 0.7%; measured change after the stop +1.92 percentage points; Lower in bothU2AF1: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyCSTB: predicted G418 readthrough 1.5%; measured change after the stop +2.20 percentage points; High in model onlyPWP2: predicted G418 readthrough 0.6%; measured change after the stop +0.71 percentage points; Lower in bothMCM3AP: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothATP5MF: predicted G418 readthrough 0.5%; measured change after the stop +0.46 percentage points; Lower in bothMAML1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothPSMC2: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothAP2M1: predicted G418 readthrough 1.1%; measured change after the stop +0.00 percentage points; High in model onlyEIF4A1: predicted G418 readthrough 0.5%; measured change after the stop +0.55 percentage points; Lower in bothPOLR3K: predicted G418 readthrough 0.8%; measured change after the stop +2.52 percentage points; Lower in bothRPL29: predicted G418 readthrough 0.3%; measured change after the stop +0.66 percentage points; Lower in bothUSP24: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothSPRTN: predicted G418 readthrough 1.3%; measured change after the stop +7.53 percentage points; High in model + measurementWDCP: predicted G418 readthrough 2.0%; measured change after the stop +2.69 percentage points; High in model onlyCHAC2: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothHNRNPDL: predicted G418 readthrough 0.3%; measured change after the stop +0.51 percentage points; Lower in bothATP1A1: predicted G418 readthrough 1.1%; measured change after the stop +0.55 percentage points; High in model onlyLRRC58: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothCCT3: predicted G418 readthrough 0.6%; measured change after the stop +0.83 percentage points; Lower in bothCIP2A: predicted G418 readthrough 0.2%; measured change after the stop +0.23 percentage points; Lower in bothWDFY3: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothTHOC7: predicted G418 readthrough 0.2%; measured change after the stop +1.49 percentage points; Lower in bothPSMD6: predicted G418 readthrough 0.3%; measured change after the stop +0.21 percentage points; Lower in bothRPL9: predicted G418 readthrough 0.3%; measured change after the stop +0.73 percentage points; Lower in bothRPN1: predicted G418 readthrough 0.9%; measured change after the stop +0.00 percentage points; High in model onlyADH5: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothH2AZ1: predicted G418 readthrough 0.3%; measured change after the stop +0.40 percentage points; Lower in bothPGRMC2: predicted G418 readthrough 1.2%; measured change after the stop +5.00 percentage points; High in model + measurementHSPA4L: predicted G418 readthrough 0.3%; measured change after the stop −0.22 percentage points; Lower in bothWDR82: predicted G418 readthrough 3.1%; measured change after the stop +0.65 percentage points; High in model onlyHMGB2: predicted G418 readthrough 0.4%; measured change after the stop −0.31 percentage points; Lower in bothSAP30: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothMAD2L1: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothNAA15: predicted G418 readthrough 0.7%; measured change after the stop +0.31 percentage points; Lower in bothICE1: predicted G418 readthrough 1.4%; measured change after the stop +1.20 percentage points; High in model onlyABCE1: predicted G418 readthrough 0.7%; measured change after the stop +1.22 percentage points; Lower in bothPRRC1: predicted G418 readthrough 0.5%; measured change after the stop +0.16 percentage points; Lower in bothRPS23: predicted G418 readthrough 0.4%; measured change after the stop +0.06 percentage points; Lower in bothMAP1B: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothNPM1: predicted G418 readthrough 0.5%; measured change after the stop +0.29 percentage points; Lower in bothFABP5: predicted G418 readthrough 0.4%; measured change after the stop +2.36 percentage points; Lower in bothRAD21: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothFMC1: predicted G418 readthrough 0.5%; measured change after the stop +4.10 percentage points; Lower in bothPHAX: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothSLC25A32: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothVIRMA: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothNOL6: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothFBXO33: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothSPTSSA: predicted G418 readthrough 2.3%; measured change after the stop +20.39 percentage points; High in model + measurementCFL2: predicted G418 readthrough 0.5%; measured change after the stop +0.19 percentage points; Lower in bothRPL36AL: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothDNAAF2: predicted G418 readthrough 0.5%; measured change after the stop +1.04 percentage points; Lower in bothPRDX3: predicted G418 readthrough 0.3%; measured change after the stop +0.56 percentage points; Lower in bothHPRT1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothGAS1: predicted G418 readthrough 1.4%; measured change after the stop +13.70 percentage points; High in model + measurementPSMC3: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothCPSF2: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothVPS35: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothIKBIP: predicted G418 readthrough 0.5%; measured change after the stop −0.43 percentage points; Lower in bothNDUFB8: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothHPS6: predicted G418 readthrough 3.4%; measured change after the stop +5.65 percentage points; High in model + measurementCCT2: predicted G418 readthrough 0.4%; measured change after the stop +0.18 percentage points; Lower in bothRNF169: predicted G418 readthrough 0.8%; measured change after the stop +1.59 percentage points; Lower in bothTMED3: predicted G418 readthrough 1.3%; measured change after the stop +5.58 percentage points; High in model + measurementHSP90B1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothPPIB: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothTERF2IP: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothMAP1A: predicted G418 readthrough 1.2%; measured change after the stop +0.00 percentage points; High in model onlyMAPRE2: predicted G418 readthrough 2.3%; measured change after the stop +9.64 percentage points; High in model + measurementPDIA3: predicted G418 readthrough 0.5%; measured change after the stop +0.41 percentage points; Lower in bothNUDT21: predicted G418 readthrough 0.6%; measured change after the stop +1.07 percentage points; Lower in bothPHB1: predicted G418 readthrough 0.5%; measured change after the stop +2.48 percentage points; Lower in bothSNRPD1: predicted G418 readthrough 0.4%; measured change after the stop −0.09 percentage points; Lower in bothGCN1: predicted G418 readthrough 0.6%; measured change after the stop +0.39 percentage points; Lower in bothMED1: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothATP5MG: predicted G418 readthrough 0.6%; measured change after the stop +0.45 percentage points; Lower in bothRRM1: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothSMG8: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothCDT1: predicted G418 readthrough 0.6%; measured change after the stop +0.48 percentage points; Lower in bothKMT2D: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothTUBA1A: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothTUBA1C: predicted G418 readthrough 0.6%; measured change after the stop +0.77 percentage points; Lower in bothTSR1: predicted G418 readthrough 0.3%; measured change after the stop +0.21 percentage points; Lower in bothPRDX2: predicted G418 readthrough 0.6%; measured change after the stop +0.06 percentage points; Lower in bothATP5PD: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothTK1: predicted G418 readthrough 0.6%; measured change after the stop +3.01 percentage points; Lower in bothECI1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothKCTD5: predicted G418 readthrough 0.5%; measured change after the stop +1.00 percentage points; Lower in bothSRRM2: predicted G418 readthrough 0.2%; measured change after the stop +0.07 percentage points; Lower in bothDDB1: predicted G418 readthrough 1.2%; measured change after the stop +1.75 percentage points; High in model onlyHNRNPUL2: predicted G418 readthrough 0.6%; measured change after the stop +0.97 percentage points; Lower in bothRPSA: predicted G418 readthrough 0.4%; measured change after the stop +0.56 percentage points; Lower in bothIRF2BP1: predicted G418 readthrough 0.4%; measured change after the stop +1.02 percentage points; Lower in bothUBB: predicted G418 readthrough 0.4%; measured change after the stop +1.83 percentage points; Lower in bothHNRNPR: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothWDR48: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothGABARAP: predicted G418 readthrough 0.7%; measured change after the stop +1.34 percentage points; Lower in bothKIF5B: predicted G418 readthrough 0.1%; measured change after the stop +0.43 percentage points; Lower in bothTMEM256: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; High in model onlyKARS1: predicted G418 readthrough 0.4%; measured change after the stop +1.42 percentage points; Lower in bothCKAP2L: predicted G418 readthrough 0.3%; measured change after the stop +0.67 percentage points; Lower in bothCDC25A: predicted G418 readthrough 0.9%; measured change after the stop +9.09 percentage points; High in model + measurementSF3B3: predicted G418 readthrough 1.0%; measured change after the stop +0.75 percentage points; High in model onlyRGPD8: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothPDHB: predicted G418 readthrough 0.4%; measured change after the stop +3.67 percentage points; Lower in bothBUB1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothRPS9: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothCEBPB: predicted G418 readthrough 1.0%; measured change after the stop +0.74 percentage points; High in model onlyLMAN2: predicted G418 readthrough 0.7%; measured change after the stop +1.06 percentage points; Lower in bothPA2G4: predicted G418 readthrough 0.8%; measured change after the stop +1.53 percentage points; Lower in bothMMADHC: predicted G418 readthrough 1.4%; measured change after the stop +1.40 percentage points; High in model onlySHCBP1: predicted G418 readthrough 3.4%; measured change after the stop +0.55 percentage points; High in model onlyARF4: predicted G418 readthrough 0.4%; measured change after the stop +1.23 percentage points; Lower in bothPCBP1: predicted G418 readthrough 0.7%; measured change after the stop +0.73 percentage points; Lower in bothTHAP11: predicted G418 readthrough 0.7%; measured change after the stop +3.52 percentage points; Lower in bothMCM7: predicted G418 readthrough 1.0%; measured change after the stop +1.69 percentage points; High in model onlyH2AC8: predicted G418 readthrough 0.7%; measured change after the stop +4.73 percentage points; Lower in bothGPRIN1: predicted G418 readthrough 0.7%; measured change after the stop −0.63 percentage points; Lower in bothHINT1: predicted G418 readthrough 0.3%; measured change after the stop −0.59 percentage points; Lower in bothH1-4: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothATP5ME: predicted G418 readthrough 0.7%; measured change after the stop +0.40 percentage points; Lower in bothARL6IP1: predicted G418 readthrough 1.1%; measured change after the stop −0.91 percentage points; High in model onlyRRM2: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothC1GALT1C1: predicted G418 readthrough 0.9%; measured change after the stop +0.46 percentage points; High in model onlySRP9: predicted G418 readthrough 0.5%; measured change after the stop +0.83 percentage points; Lower in bothHSPA4: predicted G418 readthrough 1.6%; measured change after the stop −0.45 percentage points; High in model onlyUQCRFS1: predicted G418 readthrough 0.3%; measured change after the stop +1.13 percentage points; Lower in bothPRPF8: predicted G418 readthrough 2.8%; measured change after the stop +0.43 percentage points; High in model onlyZNHIT1: predicted G418 readthrough 0.4%; measured change after the stop +4.81 percentage points; Lower in bothMGAT2: predicted G418 readthrough 0.6%; measured change after the stop +2.53 percentage points; Lower in bothLCMT2: predicted G418 readthrough 0.2%; measured change after the stop −0.65 percentage points; Lower in bothCERS6: predicted G418 readthrough 0.8%; measured change after the stop +0.72 percentage points; Lower in bothFEN1: predicted G418 readthrough 0.4%; measured change after the stop +0.65 percentage points; Lower in bothRNF187: predicted G418 readthrough 1.0%; measured change after the stop +2.52 percentage points; High in model onlyMMGT1: predicted G418 readthrough 0.3%; measured change after the stop +1.94 percentage points; Lower in bothSCAND1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothLSM3: predicted G418 readthrough 0.7%; measured change after the stop +5.21 percentage points; High in measurement onlyMRPL13: predicted G418 readthrough 0.4%; measured change after the stop +0.62 percentage points; Lower in bothCHST14: predicted G418 readthrough 1.6%; measured change after the stop +4.59 percentage points; High in model onlyBPTF: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothMAT2A: predicted G418 readthrough 1.0%; measured change after the stop +0.78 percentage points; High in model onlyUBE2E1: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothFASN: predicted G418 readthrough 0.4%; measured change after the stop +0.35 percentage points; Lower in bothRPTOR: predicted G418 readthrough 0.6%; measured change after the stop +0.85 percentage points; Lower in bothFEM1B: predicted G418 readthrough 0.5%; measured change after the stop −1.41 percentage points; Lower in bothMPLKIP: predicted G418 readthrough 0.9%; measured change after the stop +0.00 percentage points; High in model onlyPDCD6IP: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothCCDC8: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothYWHAG: predicted G418 readthrough 0.4%; measured change after the stop +3.74 percentage points; Lower in bothSEC23A: predicted G418 readthrough 0.9%; measured change after the stop +0.00 percentage points; High in model onlyRPL15: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothSRP68: predicted G418 readthrough 0.9%; measured change after the stop +0.18 percentage points; High in model onlyRPL4: predicted G418 readthrough 0.4%; measured change after the stop +0.25 percentage points; Lower in bothCFL1: predicted G418 readthrough 0.6%; measured change after the stop +4.36 percentage points; Lower in bothPCDHGC3: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothPPM1E: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothRSF1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothDHX37: predicted G418 readthrough 2.0%; measured change after the stop +0.00 percentage points; High in model onlyALG10B: predicted G418 readthrough 0.4%; measured change after the stop +1.64 percentage points; Lower in bothU2AF2: predicted G418 readthrough 0.3%; measured change after the stop +0.28 percentage points; Lower in bothNDUFA11: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothCKS1B: predicted G418 readthrough 0.7%; measured change after the stop +4.05 percentage points; Lower in bothEEF1A1: predicted G418 readthrough 0.8%; measured change after the stop +2.41 percentage points; High in model onlyEEF2: predicted G418 readthrough 0.3%; measured change after the stop +0.34 percentage points; Lower in bothSUSD5: predicted G418 readthrough 0.3%; measured change after the stop +0.45 percentage points; Lower in bothHSPA6: predicted G418 readthrough 1.1%; measured change after the stop +0.00 percentage points; High in model onlyGGA2: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothTRMT10C: predicted G418 readthrough 0.3%; measured change after the stop −0.14 percentage points; Lower in bothSP3: predicted G418 readthrough 0.3%; measured change after the stop −0.61 percentage points; Lower in bothHLTF: predicted G418 readthrough 0.5%; measured change after the stop −0.18 percentage points; Lower in bothPSMD2: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothRBM14: predicted G418 readthrough 0.5%; measured change after the stop +0.96 percentage points; Lower in bothZCCHC12: predicted G418 readthrough 0.3%; measured change after the stop +0.56 percentage points; Lower in bothVCPIP1: predicted G418 readthrough 1.4%; measured change after the stop +0.00 percentage points; High in model onlyHCFC1: predicted G418 readthrough 0.5%; measured change after the stop +0.26 percentage points; Lower in bothCASP2: predicted G418 readthrough 0.9%; measured change after the stop +7.16 percentage points; High in model + measurementMEPCE: predicted G418 readthrough 0.3%; measured change after the stop +0.52 percentage points; Lower in bothCD164: predicted G418 readthrough 0.3%; measured change after the stop +0.77 percentage points; Lower in bothCBX2: predicted G418 readthrough 0.7%; measured change after the stop +0.13 percentage points; Lower in bothCDC20: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothRPL38: predicted G418 readthrough 0.8%; measured change after the stop +1.11 percentage points; Lower in bothHOXB9: predicted G418 readthrough 0.2%; measured change after the stop +0.52 percentage points; Lower in bothPDE12: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothB4GAT1: predicted G418 readthrough 0.7%; measured change after the stop +2.30 percentage points; Lower in bothRNF26: predicted G418 readthrough 1.1%; measured change after the stop +0.86 percentage points; High in model onlyRAB1B: predicted G418 readthrough 0.5%; measured change after the stop +1.91 percentage points; Lower in bothRPL13: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothUQCRH: predicted G418 readthrough 0.4%; measured change after the stop +2.82 percentage points; Lower in bothTRIB1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothBANF1: predicted G418 readthrough 1.2%; measured change after the stop +14.00 percentage points; High in model + measurementTMEM70: predicted G418 readthrough 0.7%; measured change after the stop +2.52 percentage points; Lower in bothCCDC85B: predicted G418 readthrough 1.0%; measured change after the stop +5.33 percentage points; High in model + measurementTAF7: predicted G418 readthrough 0.3%; measured change after the stop +0.24 percentage points; Lower in bothTUFM: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothMRPS23: predicted G418 readthrough 0.7%; measured change after the stop +0.60 percentage points; Lower in bothHNRNPD: predicted G418 readthrough 0.3%; measured change after the stop +1.14 percentage points; Lower in bothCOX8A: predicted G418 readthrough 0.5%; measured change after the stop +6.54 percentage points; High in measurement onlyRPL27A: predicted G418 readthrough 1.0%; measured change after the stop +0.56 percentage points; High in model onlyPRKDC: predicted G418 readthrough 0.5%; measured change after the stop +0.09 percentage points; Lower in bothCKS2: predicted G418 readthrough 0.7%; measured change after the stop +7.90 percentage points; High in measurement onlyKDM3B: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothPOLR2B: predicted G418 readthrough 1.3%; measured change after the stop +0.00 percentage points; High in model onlyFARSA: predicted G418 readthrough 2.4%; measured change after the stop −0.48 percentage points; High in model onlySUPT6H: predicted G418 readthrough 0.4%; measured change after the stop +0.85 percentage points; Lower in bothPFAS: predicted G418 readthrough 1.6%; measured change after the stop +0.74 percentage points; High in model onlyTLN1: predicted G418 readthrough 0.3%; measured change after the stop +0.14 percentage points; Lower in bothHNRNPA0: predicted G418 readthrough 0.5%; measured change after the stop +0.96 percentage points; Lower in bothFZD2: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothNCAPD2: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothMDH2: predicted G418 readthrough 0.6%; measured change after the stop +0.73 percentage points; Lower in bothYOD1: predicted G418 readthrough 1.1%; measured change after the stop +0.45 percentage points; High in model onlyWNK1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothRPS4X: predicted G418 readthrough 0.5%; measured change after the stop +0.88 percentage points; Lower in bothCALR: predicted G418 readthrough 0.4%; measured change after the stop +1.14 percentage points; Lower in bothHNRNPH2: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothALX1: predicted G418 readthrough 0.6%; measured change after the stop +0.81 percentage points; Lower in bothPNMA1: predicted G418 readthrough 1.1%; measured change after the stop +1.44 percentage points; High in model onlyTRIM25: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothGADD45GIP1: predicted G418 readthrough 1.0%; measured change after the stop +0.20 percentage points; High in model onlyDNAJA2: predicted G418 readthrough 0.2%; measured change after the stop +0.52 percentage points; Lower in bothLRCH2: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothCNOT1: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothMRPS24: predicted G418 readthrough 0.6%; measured change after the stop +1.08 percentage points; Lower in bothSLC25A5: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothMAGEF1: predicted G418 readthrough 0.5%; measured change after the stop +0.15 percentage points; Lower in bothUBE2N: predicted G418 readthrough 0.3%; measured change after the stop −0.73 percentage points; Lower in bothCLSPN: predicted G418 readthrough 1.3%; measured change after the stop +1.03 percentage points; High in model onlyGINS3: predicted G418 readthrough 1.1%; measured change after the stop +8.04 percentage points; High in model + measurementPABPC1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothCYC1: predicted G418 readthrough 2.0%; measured change after the stop +4.27 percentage points; High in model onlyGEMIN4: predicted G418 readthrough 2.0%; measured change after the stop +1.71 percentage points; High in model onlyTALDO1: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; High in model onlyKLHL11: predicted G418 readthrough 0.4%; measured change after the stop +0.53 percentage points; Lower in bothF2R: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothPRDX1: predicted G418 readthrough 0.4%; measured change after the stop +6.82 percentage points; High in measurement onlyDCTPP1: predicted G418 readthrough 0.4%; measured change after the stop +1.53 percentage points; Lower in bothUBE2O: predicted G418 readthrough 0.9%; measured change after the stop +0.00 percentage points; High in model onlyRBBP6: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothHSPH1: predicted G418 readthrough 0.5%; measured change after the stop +1.52 percentage points; Lower in bothRNPS1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothPAGR1: predicted G418 readthrough 1.1%; measured change after the stop +3.32 percentage points; High in model onlyZNF121: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothPOLE: predicted G418 readthrough 1.2%; measured change after the stop +0.00 percentage points; High in model onlyCHORDC1: predicted G418 readthrough 1.1%; measured change after the stop +0.00 percentage points; High in model onlyRBMX: predicted G418 readthrough 0.4%; measured change after the stop +1.20 percentage points; Lower in bothBLOC1S4: predicted G418 readthrough 2.7%; measured change after the stop +4.20 percentage points; High in model onlyMRFAP1L1: predicted G418 readthrough 0.9%; measured change after the stop +3.87 percentage points; High in model onlySMCHD1: predicted G418 readthrough 0.8%; measured change after the stop +0.96 percentage points; High in model onlyTOMM5: predicted G418 readthrough 0.3%; measured change after the stop +2.86 percentage points; Lower in bothSENP3: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothYBX1: predicted G418 readthrough 0.3%; measured change after the stop +0.20 percentage points; Lower in bothTCP1: predicted G418 readthrough 1.5%; measured change after the stop +0.86 percentage points; High in model onlySFXN1: predicted G418 readthrough 0.3%; measured change after the stop +1.22 percentage points; Lower in bothPOLR2L: predicted G418 readthrough 1.7%; measured change after the stop +2.19 percentage points; High in model onlySET: predicted G418 readthrough 0.5%; measured change after the stop +0.18 percentage points; Lower in bothPPP2R1A: predicted G418 readthrough 2.0%; measured change after the stop +3.17 percentage points; High in model onlyNPEPPS: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothRPP25: predicted G418 readthrough 1.3%; measured change after the stop +0.46 percentage points; High in model onlySMC1A: predicted G418 readthrough 0.7%; measured change after the stop +0.58 percentage points; Lower in bothUBA6: predicted G418 readthrough 0.8%; measured change after the stop +0.88 percentage points; Lower in bothWDR33: predicted G418 readthrough 0.4%; measured change after the stop +1.62 percentage points; Lower in bothCOX5A: predicted G418 readthrough 0.2%; measured change after the stop +1.76 percentage points; Lower in bothEEA1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothSF3B2: predicted G418 readthrough 0.7%; measured change after the stop +0.55 percentage points; Lower in bothRUVBL1: predicted G418 readthrough 0.6%; measured change after the stop +0.69 percentage points; Lower in bothNCL: predicted G418 readthrough 0.9%; measured change after the stop +0.08 percentage points; High in model onlyMAZ: predicted G418 readthrough 0.5%; measured change after the stop +0.91 percentage points; Lower in bothRPS15A: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothTYMS: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothRPL7A: predicted G418 readthrough 0.4%; measured change after the stop +0.22 percentage points; Lower in bothPOGLUT3: predicted G418 readthrough 1.1%; measured change after the stop +3.81 percentage points; High in model onlyRAP2B: predicted G418 readthrough 1.0%; measured change after the stop +1.56 percentage points; High in model onlyMETAP2: predicted G418 readthrough 0.4%; measured change after the stop +0.67 percentage points; Lower in bothDEGS1: predicted G418 readthrough 0.3%; measured change after the stop −0.24 percentage points; Lower in bothNDRG1: predicted G418 readthrough 0.4%; measured change after the stop +0.98 percentage points; Lower in bothSNRNP200: predicted G418 readthrough 1.0%; measured change after the stop +0.15 percentage points; High in model onlyCSTF3: predicted G418 readthrough 0.3%; measured change after the stop +0.63 percentage points; Lower in bothMAP1S: predicted G418 readthrough 1.3%; measured change after the stop +0.00 percentage points; High in model onlyPRMT5: predicted G418 readthrough 1.4%; measured change after the stop +1.55 percentage points; High in model onlyHSPA5: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothRNF40: predicted G418 readthrough 1.2%; measured change after the stop +0.99 percentage points; High in model onlyMED14: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothARID1A: predicted G418 readthrough 2.2%; measured change after the stop +1.65 percentage points; High in model onlyHMGCS1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothLMNB2: predicted G418 readthrough 0.7%; measured change after the stop +0.69 percentage points; Lower in bothSOX2: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothHASPIN: predicted G418 readthrough 0.2%; measured change after the stop −0.49 percentage points; Lower in bothHNRNPM: predicted G418 readthrough 0.4%; measured change after the stop +0.02 percentage points; Lower in bothYLPM1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothABCF1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothITM2C: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothMAGED1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothIMPDH2: predicted G418 readthrough 1.2%; measured change after the stop +0.00 percentage points; High in model onlyNDUFAF3: predicted G418 readthrough 0.8%; measured change after the stop +1.23 percentage points; High in model onlyCARM1: predicted G418 readthrough 0.8%; measured change after the stop +0.88 percentage points; Lower in bothPAK2: predicted G418 readthrough 0.7%; measured change after the stop +2.24 percentage points; Lower in bothTIMM22: predicted G418 readthrough 0.6%; measured change after the stop +2.07 percentage points; Lower in bothKHDRBS1: predicted G418 readthrough 0.5%; measured change after the stop −0.40 percentage points; Lower in bothPRR14L: predicted G418 readthrough 0.9%; measured change after the stop +0.48 percentage points; High in model onlyADI1: predicted G418 readthrough 1.4%; measured change after the stop +2.65 percentage points; High in model onlySP1: predicted G418 readthrough 1.0%; measured change after the stop +2.02 percentage points; High in model onlyNDUFA12: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothTUBB: predicted G418 readthrough 0.4%; measured change after the stop +0.02 percentage points; Lower in bothCOL4A5: predicted G418 readthrough 0.5%; measured change after the stop +0.96 percentage points; Lower in bothPOU3F2: predicted G418 readthrough 0.6%; measured change after the stop +0.64 percentage points; Lower in bothPTRHD1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothCOA3: predicted G418 readthrough 0.5%; measured change after the stop +3.25 percentage points; Lower in bothTYW2: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothNOP10: predicted G418 readthrough 0.9%; measured change after the stop +18.25 percentage points; High in model + measurementFOXK1: predicted G418 readthrough 0.4%; measured change after the stop +3.21 percentage points; Lower in bothEEF1G: predicted G418 readthrough 0.6%; measured change after the stop +0.19 percentage points; Lower in bothPDLIM1: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothMARCKSL1: predicted G418 readthrough 0.9%; measured change after the stop +8.19 percentage points; High in model + measurementRGPD6: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothPYCR1: predicted G418 readthrough 3.2%; measured change after the stop +6.95 percentage points; High in model + measurementUQCR10: predicted G418 readthrough 0.3%; measured change after the stop +1.93 percentage points; Lower in bothBRCC3: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothMRPL54: predicted G418 readthrough 0.9%; measured change after the stop +3.19 percentage points; High in model onlyMRTO4: predicted G418 readthrough 1.1%; measured change after the stop +0.24 percentage points; High in model onlyVMA21: predicted G418 readthrough 0.4%; measured change after the stop +3.63 percentage points; Lower in bothCREB3L2: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothATP6V0C: predicted G418 readthrough 0.5%; measured change after the stop +1.90 percentage points; Lower in bothKPNA2: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothINTS5: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothKDELR1: predicted G418 readthrough 1.1%; measured change after the stop +4.76 percentage points; High in model onlyDNAJA1: predicted G418 readthrough 0.5%; measured change after the stop +2.48 percentage points; Lower in bothH2AC21: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothNPLOC4: predicted G418 readthrough 0.6%; measured change after the stop +1.48 percentage points; Lower in bothCSTF2T: predicted G418 readthrough 1.1%; measured change after the stop +0.65 percentage points; High in model onlyDUT: predicted G418 readthrough 0.5%; measured change after the stop +1.40 percentage points; Lower in bothPURA: predicted G418 readthrough 1.5%; measured change after the stop +1.07 percentage points; High in model onlyGLRX5: predicted G418 readthrough 0.5%; measured change after the stop +0.02 percentage points; Lower in bothH1-5: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothDRG1: predicted G418 readthrough 0.8%; measured change after the stop +5.37 percentage points; High in measurement onlyZNF703: predicted G418 readthrough 0.6%; measured change after the stop +0.38 percentage points; Lower in bothEIF3C: predicted G418 readthrough 0.5%; measured change after the stop +0.35 percentage points; Lower in bothSELENOF: predicted G418 readthrough 0.2%; measured change after the stop +1.77 percentage points; Lower in bothGLUL: predicted G418 readthrough 0.5%; measured change after the stop +0.72 percentage points; Lower in bothVDAC2: predicted G418 readthrough 0.6%; measured change after the stop +0.91 percentage points; Lower in bothC6orf120: predicted G418 readthrough 1.0%; measured change after the stop +3.22 percentage points; High in model onlyNSMCE3: predicted G418 readthrough 1.1%; measured change after the stop +9.33 percentage points; High in model + measurementEMILIN3: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothDDX28: predicted G418 readthrough 1.0%; measured change after the stop −0.14 percentage points; High in model onlyHEXIM1: predicted G418 readthrough 0.6%; measured change after the stop +0.79 percentage points; Lower in bothTMED9: predicted G418 readthrough 0.7%; measured change after the stop +4.18 percentage points; Lower in bothRLIM: predicted G418 readthrough 0.4%; measured change after the stop +0.60 percentage points; Lower in bothMRPL40: predicted G418 readthrough 0.3%; measured change after the stop +0.80 percentage points; Lower in bothH2AC14: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothKNTC1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothPGP: predicted G418 readthrough 0.3%; measured change after the stop −0.02 percentage points; Lower in bothBSG: predicted G418 readthrough 1.0%; measured change after the stop +4.52 percentage points; High in model onlyTUBB2A: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothMRPL12: predicted G418 readthrough 0.5%; measured change after the stop +3.41 percentage points; Lower in bothH1-10: predicted G418 readthrough 0.4%; measured change after the stop +1.95 percentage points; Lower in bothARID2: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothERCC6L: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyPGAM1: predicted G418 readthrough 0.6%; measured change after the stop +1.06 percentage points; Lower in bothRSBN1L: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothPPP1CC: predicted G418 readthrough 0.4%; measured change after the stop +0.28 percentage points; Lower in bothPKM: predicted G418 readthrough 0.9%; measured change after the stop +0.09 percentage points; High in model onlyPITPNB: predicted G418 readthrough 0.4%; measured change after the stop +0.36 percentage points; Lower in bothSF3B1: predicted G418 readthrough 0.5%; measured change after the stop +0.05 percentage points; Lower in bothATP5IF1: predicted G418 readthrough 0.5%; measured change after the stop +1.45 percentage points; Lower in bothUBA1: predicted G418 readthrough 2.3%; measured change after the stop +0.00 percentage points; High in model onlyTUBA1B: predicted G418 readthrough 0.5%; measured change after the stop +0.64 percentage points; Lower in bothSPTY2D1: predicted G418 readthrough 1.2%; measured change after the stop +1.30 percentage points; High in model onlyEIF5A: predicted G418 readthrough 0.6%; measured change after the stop +0.38 percentage points; Lower in bothBRIX1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothATXN2L: predicted G418 readthrough 0.4%; measured change after the stop −0.61 percentage points; Lower in bothDHX15: predicted G418 readthrough 0.6%; measured change after the stop +0.98 percentage points; Lower in bothMRPL35: predicted G418 readthrough 0.5%; measured change after the stop +3.20 percentage points; Lower in bothTNPO1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothCLPTM1: predicted G418 readthrough 1.3%; measured change after the stop +0.00 percentage points; High in model onlyZNF644: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothRTN4: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; High in model onlyPRP4K: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothZC3H15: predicted G418 readthrough 2.3%; measured change after the stop +0.00 percentage points; High in model onlySPRYD4: predicted G418 readthrough 0.9%; measured change after the stop +1.49 percentage points; High in model onlyMRPL30: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothAPLP2: predicted G418 readthrough 0.5%; measured change after the stop +2.56 percentage points; Lower in bothPRIM1: predicted G418 readthrough 0.8%; measured change after the stop −0.56 percentage points; Lower in bothUBQLN2: predicted G418 readthrough 0.3%; measured change after the stop +0.07 percentage points; Lower in bothAURKAIP1: predicted G418 readthrough 0.7%; measured change after the stop +0.69 percentage points; Lower in bothMTSS2: predicted G418 readthrough 1.9%; measured change after the stop +2.25 percentage points; High in model onlyEIF4EBP1: predicted G418 readthrough 0.3%; measured change after the stop +1.23 percentage points; Lower in bothNCR3LG1: predicted G418 readthrough 0.3%; measured change after the stop −0.43 percentage points; Lower in bothANP32B: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothBAZ1B: predicted G418 readthrough 0.3%; measured change after the stop −0.10 percentage points; Lower in bothCOXFA4: predicted G418 readthrough 0.6%; measured change after the stop +0.09 percentage points; Lower in bothUBC: predicted G418 readthrough 0.3%; measured change after the stop −0.04 percentage points; Lower in bothSERINC1: predicted G418 readthrough 1.0%; measured change after the stop +0.11 percentage points; High in model onlyH2BC11: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothUSP1: predicted G418 readthrough 0.6%; measured change after the stop +1.53 percentage points; Lower in bothEIF4G2: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothZNF326: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothTUBB4B: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothPRDX6: predicted G418 readthrough 0.2%; measured change after the stop +0.37 percentage points; Lower in bothH3-5: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothNUP43: predicted G418 readthrough 0.4%; measured change after the stop +0.75 percentage points; Lower in bothGSPT2: predicted G418 readthrough 0.4%; measured change after the stop +0.91 percentage points; Lower in bothRRP1B: predicted G418 readthrough 0.8%; measured change after the stop +1.36 percentage points; Lower in bothCCNG1: predicted G418 readthrough 0.6%; measured change after the stop +1.68 percentage points; Lower in bothH1-0: predicted G418 readthrough 1.9%; measured change after the stop +0.60 percentage points; High in model onlyHNRNPA1: predicted G418 readthrough 0.8%; measured change after the stop +0.20 percentage points; Lower in bothH2AC7: predicted G418 readthrough 0.3%; measured change after the stop +7.95 percentage points; High in measurement onlyRTL6: predicted G418 readthrough 0.5%; measured change after the stop +0.36 percentage points; Lower in bothCAPRIN1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothHMGB1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothTAF9B: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothUBE2L3: predicted G418 readthrough 0.4%; measured change after the stop +0.48 percentage points; Lower in bothSOX12: predicted G418 readthrough 0.6%; measured change after the stop +5.46 percentage points; High in measurement onlySNRPD2: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothMAGEH1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothNELFB: predicted G418 readthrough 0.9%; measured change after the stop +2.65 percentage points; High in model onlyDPYSL3: predicted G418 readthrough 0.6%; measured change after the stop +1.68 percentage points; Lower in bothLYAR: predicted G418 readthrough 0.7%; measured change after the stop +0.97 percentage points; Lower in bothHOXA9: predicted G418 readthrough 1.0%; measured change after the stop +3.81 percentage points; High in model onlyRPS21: predicted G418 readthrough 2.4%; measured change after the stop +6.33 percentage points; High in model + measurementPPM1G: predicted G418 readthrough 1.3%; measured change after the stop −0.41 percentage points; High in model onlyRPL28: predicted G418 readthrough 0.9%; measured change after the stop +0.43 percentage points; High in model onlyYTHDC1: predicted G418 readthrough 0.3%; measured change after the stop +0.94 percentage points; Lower in bothUSP7: predicted G418 readthrough 1.0%; measured change after the stop +0.15 percentage points; High in model onlyMRPL20: predicted G418 readthrough 0.5%; measured change after the stop +1.63 percentage points; Lower in bothBCAP31: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothLIN28B: predicted G418 readthrough 0.5%; measured change after the stop +3.29 percentage points; Lower in bothARHGAP5: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothUBE2G2: predicted G418 readthrough 0.7%; measured change after the stop +1.71 percentage points; Lower in bothCLTA: predicted G418 readthrough 0.5%; measured change after the stop +1.32 percentage points; Lower in bothEIF4G1: predicted G418 readthrough 0.6%; measured change after the stop +0.41 percentage points; Lower in bothE2F3: predicted G418 readthrough 3.0%; measured change after the stop +0.00 percentage points; High in model onlyAPP: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothRPL36: predicted G418 readthrough 0.9%; measured change after the stop +0.00 percentage points; High in model onlyHGH1: predicted G418 readthrough 0.7%; measured change after the stop +1.65 percentage points; Lower in bothRAC1: predicted G418 readthrough 0.3%; measured change after the stop −0.64 percentage points; Lower in bothHELLS: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothCKB: predicted G418 readthrough 0.7%; measured change after the stop +5.18 percentage points; High in measurement onlyWDR3: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothTBX3: predicted G418 readthrough 0.4%; measured change after the stop +2.14 percentage points; Lower in bothCTNNB1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothSAMM50: predicted G418 readthrough 0.6%; measured change after the stop +4.47 percentage points; Lower in bothNASP: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothBIRC5: predicted G418 readthrough 1.0%; measured change after the stop +11.44 percentage points; High in model + measurementSEPTIN7: predicted G418 readthrough 0.5%; measured change after the stop +0.59 percentage points; Lower in bothSEC13: predicted G418 readthrough 1.7%; measured change after the stop +7.82 percentage points; High in model + measurementATR: predicted G418 readthrough 0.7%; measured change after the stop −0.89 percentage points; Lower in bothCS: predicted G418 readthrough 0.2%; measured change after the stop +1.39 percentage points; Lower in bothACLY: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothPTTG1: predicted G418 readthrough 0.3%; measured change after the stop −1.05 percentage points; Lower in bothRPL7: predicted G418 readthrough 0.2%; measured change after the stop +0.09 percentage points; Lower in bothYWHAB: predicted G418 readthrough 0.4%; measured change after the stop +1.78 percentage points; Lower in bothDDX21: predicted G418 readthrough 0.6%; measured change after the stop +0.50 percentage points; Lower in bothNDUFV3: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothAP2A1: predicted G418 readthrough 1.0%; measured change after the stop +1.00 percentage points; High in model onlyCRKL: predicted G418 readthrough 0.9%; measured change after the stop +1.89 percentage points; High in model onlyCOPS8: predicted G418 readthrough 1.0%; measured change after the stop +6.84 percentage points; High in model + measurementTHRAP3: predicted G418 readthrough 0.4%; measured change after the stop +0.83 percentage points; Lower in bothPES1: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothALDH9A1: predicted G418 readthrough 1.2%; measured change after the stop +1.10 percentage points; High in model onlyHTT: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothIFT70A: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothPLS3: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothPRELID3B: predicted G418 readthrough 0.9%; measured change after the stop +0.00 percentage points; High in model onlyPSMD12: predicted G418 readthrough 0.4%; measured change after the stop +0.75 percentage points; Lower in bothFBRS: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothGTF3C1: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; High in model onlyG3BP1: predicted G418 readthrough 0.5%; measured change after the stop +1.94 percentage points; Lower in bothUBE2C: predicted G418 readthrough 2.2%; measured change after the stop +4.86 percentage points; High in model onlyH3C4: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothSTOML2: predicted G418 readthrough 0.9%; measured change after the stop +1.50 percentage points; High in model onlyGANAB: predicted G418 readthrough 0.7%; measured change after the stop +0.68 percentage points; Lower in bothPPP4R2: predicted G418 readthrough 1.1%; measured change after the stop +0.00 percentage points; High in model onlyBICD2: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothPTBP1: predicted G418 readthrough 0.5%; measured change after the stop +1.16 percentage points; Lower in bothH2BC12: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothIGF2R: predicted G418 readthrough 1.6%; measured change after the stop −0.18 percentage points; High in model onlyCALM1: predicted G418 readthrough 0.6%; measured change after the stop +1.27 percentage points; Lower in bothSCRIB: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothGM2A: predicted G418 readthrough 0.4%; measured change after the stop +1.50 percentage points; Lower in bothSFPQ: predicted G418 readthrough 0.6%; measured change after the stop −0.09 percentage points; Lower in bothHDGF: predicted G418 readthrough 0.7%; measured change after the stop +7.45 percentage points; High in measurement onlyLAGE3: predicted G418 readthrough 0.3%; measured change after the stop +1.45 percentage points; Lower in bothSMC4: predicted G418 readthrough 1.1%; measured change after the stop +0.33 percentage points; High in model onlyHNRNPA1L2: predicted G418 readthrough 0.8%; measured change after the stop +0.37 percentage points; Lower in bothAP1B1: predicted G418 readthrough 0.7%; measured change after the stop +0.97 percentage points; Lower in bothSGO2: predicted G418 readthrough 0.4%; measured change after the stop +0.46 percentage points; Lower in bothDDX47: predicted G418 readthrough 0.4%; measured change after the stop −0.65 percentage points; Lower in bothRAD23B: predicted G418 readthrough 1.2%; measured change after the stop +4.28 percentage points; High in model onlyDMAC1: predicted G418 readthrough 0.9%; measured change after the stop +1.98 percentage points; High in model onlyHNRNPAB: predicted G418 readthrough 0.7%; measured change after the stop +1.79 percentage points; Lower in bothTOMM7: predicted G418 readthrough 0.4%; measured change after the stop −0.22 percentage points; Lower in bothZXDA: predicted G418 readthrough 0.9%; measured change after the stop +0.94 percentage points; High in model onlyQSOX2: predicted G418 readthrough 0.5%; measured change after the stop +4.37 percentage points; Lower in bothBDP1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothH2AC13: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothVCP: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothKLHL9: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothH2AC11: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothH3C12: predicted G418 readthrough 0.5%; measured change after the stop +0.94 percentage points; Lower in bothDNMT1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothH2AC17: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothARPC5: predicted G418 readthrough 0.2%; measured change after the stop +0.57 percentage points; Lower in bothATG2B: predicted G418 readthrough 1.7%; measured change after the stop −0.30 percentage points; High in model onlySRSF2: predicted G418 readthrough 0.4%; measured change after the stop +2.50 percentage points; Lower in bothXRCC6: predicted G418 readthrough 2.7%; measured change after the stop +2.72 percentage points; High in model onlyTFRC: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothVPS13A: predicted G418 readthrough 3.2%; measured change after the stop +0.00 percentage points; High in model onlyBAZ1A: predicted G418 readthrough 1.3%; measured change after the stop +2.39 percentage points; High in model onlyZNF460: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothDSC3: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothETF1: predicted G418 readthrough 0.5%; measured change after the stop +1.12 percentage points; Lower in bothSCAF1: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyANKHD1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothMRPL37: predicted G418 readthrough 0.6%; measured change after the stop −0.72 percentage points; Lower in bothEIF3B: predicted G418 readthrough 1.2%; measured change after the stop +3.71 percentage points; High in model onlyMPC1: predicted G418 readthrough 0.9%; measured change after the stop +5.04 percentage points; High in model + measurementGSTM3: predicted G418 readthrough 0.6%; measured change after the stop −0.62 percentage points; Lower in bothSERBP1: predicted G418 readthrough 0.8%; measured change after the stop +1.52 percentage points; Lower in bothBEX3: predicted G418 readthrough 1.7%; measured change after the stop +2.07 percentage points; High in model onlyTOP1: predicted G418 readthrough 1.2%; measured change after the stop +0.20 percentage points; High in model onlyCTR9: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothLTN1: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyHMGN2: predicted G418 readthrough 0.6%; measured change after the stop +9.95 percentage points; High in measurement onlyZNF318: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothAPOBEC3C: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothMTOR: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothMAGEE1: predicted G418 readthrough 0.3%; measured change after the stop +1.42 percentage points; Lower in bothPNP: predicted G418 readthrough 3.6%; measured change after the stop +4.44 percentage points; High in model onlyACOT7: predicted G418 readthrough 0.4%; measured change after the stop +1.60 percentage points; Lower in bothOSTC: predicted G418 readthrough 0.4%; measured change after the stop +1.77 percentage points; Lower in bothRPL39: predicted G418 readthrough 0.4%; measured change after the stop +2.11 percentage points; Lower in bothPSMB5: predicted G418 readthrough 0.8%; measured change after the stop +2.79 percentage points; Lower in bothSMC3: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothCOL6A1: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothMSRB1: predicted G418 readthrough 0.3%; measured change after the stop +1.15 percentage points; Lower in bothILF2: predicted G418 readthrough 1.4%; measured change after the stop +0.73 percentage points; High in model onlyZNF830: predicted G418 readthrough 0.6%; measured change after the stop +0.34 percentage points; Lower in bothCHML: predicted G418 readthrough 0.6%; measured change after the stop +3.13 percentage points; Lower in bothMTR: predicted G418 readthrough 0.5%; measured change after the stop +1.25 percentage points; Lower in bothHEATR1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothTOMM20: predicted G418 readthrough 0.7%; measured change after the stop +2.70 percentage points; Lower in bothIRF2BP2: predicted G418 readthrough 1.3%; measured change after the stop +1.85 percentage points; High in model onlyCOA6: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothEXOC8: predicted G418 readthrough 0.4%; measured change after the stop +1.24 percentage points; Lower in bothH3-4: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothIBA57: predicted G418 readthrough 0.3%; measured change after the stop +1.39 percentage points; Lower in bothPARP1: predicted G418 readthrough 0.3%; measured change after the stop +0.19 percentage points; Lower in bothACBD3: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothH3-3A: predicted G418 readthrough 0.4%; measured change after the stop −0.60 percentage points; Lower in bothFBXO28: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothIARS2: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothEPRS1: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothRRP15: predicted G418 readthrough 0.3%; measured change after the stop +0.68 percentage points; Lower in bothCENPF: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothDTL: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothSLC30A1: predicted G418 readthrough 0.6%; measured change after the stop +0.58 percentage points; Lower in bothMRPL18: predicted G418 readthrough 0.4%; measured change after the stop −0.01 percentage points; Lower in bothACAT2: predicted G418 readthrough 0.7%; measured change after the stop +0.67 percentage points; Lower in bothDYNLT1: predicted G418 readthrough 1.7%; measured change after the stop +5.28 percentage points; High in model + measurementNUCKS1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothSCAF8: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothPPP1R15B: predicted G418 readthrough 0.9%; measured change after the stop +1.21 percentage points; High in model onlyRABIF: predicted G418 readthrough 0.4%; measured change after the stop +1.30 percentage points; Lower in bothTIMM17A: predicted G418 readthrough 0.4%; measured change after the stop +0.74 percentage points; Lower in bothDCPH1: predicted G418 readthrough 2.2%; measured change after the stop +2.82 percentage points; High in model onlyMTHFD1L: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothTMEM9: predicted G418 readthrough 0.7%; measured change after the stop +7.84 percentage points; High in measurement onlyKIF14: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothZNF281: predicted G418 readthrough 0.2%; measured change after the stop −0.46 percentage points; Lower in bothTPR: predicted G418 readthrough 0.5%; measured change after the stop +0.06 percentage points; Lower in bothDHX9: predicted G418 readthrough 0.4%; measured change after the stop −0.16 percentage points; Lower in bothSF3B5: predicted G418 readthrough 0.7%; measured change after the stop +10.63 percentage points; High in measurement onlyLTV1: predicted G418 readthrough 0.5%; measured change after the stop +1.51 percentage points; Lower in bothIER5: predicted G418 readthrough 1.0%; measured change after the stop +6.57 percentage points; High in model + measurementSOAT1: predicted G418 readthrough 0.7%; measured change after the stop +0.93 percentage points; Lower in bothVTA1: predicted G418 readthrough 0.7%; measured change after the stop +0.05 percentage points; Lower in bothCITED2: predicted G418 readthrough 2.3%; measured change after the stop +12.90 percentage points; High in model + measurementCACYBP: predicted G418 readthrough 1.1%; measured change after the stop +0.75 percentage points; High in model onlyTIPRL: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothUCK2: predicted G418 readthrough 3.6%; measured change after the stop +3.23 percentage points; High in model onlySDHC: predicted G418 readthrough 0.5%; measured change after the stop +2.68 percentage points; Lower in bothUFC1: predicted G418 readthrough 0.6%; measured change after the stop +1.26 percentage points; Lower in bothPFDN2: predicted G418 readthrough 0.7%; measured change after the stop +0.80 percentage points; Lower in bothPIGM: predicted G418 readthrough 0.6%; measured change after the stop +0.71 percentage points; Lower in bothTAGLN2: predicted G418 readthrough 1.5%; measured change after the stop +0.49 percentage points; High in model onlyETV3: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothCENPW: predicted G418 readthrough 0.4%; measured change after the stop +0.31 percentage points; Lower in bothMTX1: predicted G418 readthrough 1.1%; measured change after the stop +0.00 percentage points; High in model onlyMAN1A1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothTSPYL1: predicted G418 readthrough 0.6%; measured change after the stop +3.23 percentage points; Lower in bothMKI67: predicted G418 readthrough 2.1%; measured change after the stop +0.00 percentage points; High in model onlyCHTOP: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyS100A10: predicted G418 readthrough 0.3%; measured change after the stop +3.32 percentage points; Lower in bothOAT: predicted G418 readthrough 0.9%; measured change after the stop +0.77 percentage points; High in model onlyBUB3: predicted G418 readthrough 2.2%; measured change after the stop +3.63 percentage points; High in model onlyPSMD4: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothVPS72: predicted G418 readthrough 0.6%; measured change after the stop +2.07 percentage points; Lower in bothLYSMD1: predicted G418 readthrough 1.5%; measured change after the stop +1.69 percentage points; High in model onlyMCL1: predicted G418 readthrough 0.5%; measured change after the stop +1.05 percentage points; Lower in bothRPRD2: predicted G418 readthrough 1.3%; measured change after the stop +0.87 percentage points; High in model onlyMCMBP: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothBAG3: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothEIF3A: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothH2BC21: predicted G418 readthrough 0.6%; measured change after the stop +7.30 percentage points; High in measurement onlyH3C14: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothH2AC18: predicted G418 readthrough 0.6%; measured change after the stop +0.97 percentage points; Lower in bothASCC3: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothH2BC18: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothNHLRC2: predicted G418 readthrough 1.2%; measured change after the stop +0.00 percentage points; High in model onlySCAF11: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothMDN1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothNRAS: predicted G418 readthrough 0.7%; measured change after the stop +0.98 percentage points; Lower in bothBCAS2: predicted G418 readthrough 1.2%; measured change after the stop +4.08 percentage points; High in model onlySYNCRIP: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothSLC16A1: predicted G418 readthrough 0.7%; measured change after the stop +0.59 percentage points; Lower in bothGSTO1: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyATP5PB: predicted G418 readthrough 0.3%; measured change after the stop +0.22 percentage points; Lower in bothSLK: predicted G418 readthrough 0.8%; measured change after the stop +1.54 percentage points; Lower in bothATP6AP1: predicted G418 readthrough 1.6%; measured change after the stop +2.26 percentage points; High in model onlyRBM15: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothTTK: predicted G418 readthrough 1.1%; measured change after the stop +0.00 percentage points; High in model onlyELOVL4: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothRPL10: predicted G418 readthrough 0.6%; measured change after the stop +1.09 percentage points; Lower in bothFLNA: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothGNAI3: predicted G418 readthrough 0.8%; measured change after the stop +0.51 percentage points; High in model onlyPRMT6: predicted G418 readthrough 0.7%; measured change after the stop +1.17 percentage points; Lower in bothSSR4: predicted G418 readthrough 0.9%; measured change after the stop +0.00 percentage points; High in model onlyCNN3: predicted G418 readthrough 0.5%; measured change after the stop +2.93 percentage points; Lower in bothCETN2: predicted G418 readthrough 0.3%; measured change after the stop +1.36 percentage points; Lower in bothMAGEA3: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothRPL5: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothYTHDF1: predicted G418 readthrough 0.7%; measured change after the stop +1.76 percentage points; Lower in bothSCD: predicted G418 readthrough 0.6%; measured change after the stop +0.88 percentage points; Lower in bothLRRC8C: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothGTF2B: predicted G418 readthrough 0.4%; measured change after the stop +1.72 percentage points; Lower in bothLDOC1: predicted G418 readthrough 0.6%; measured change after the stop +5.21 percentage points; High in measurement onlyZNHIT6: predicted G418 readthrough 0.8%; measured change after the stop +1.09 percentage points; Lower in bothRPF1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothFKBP1C: predicted G418 readthrough 2.2%; measured change after the stop +4.91 percentage points; High in model onlyRTL8A: predicted G418 readthrough 0.7%; measured change after the stop +1.63 percentage points; Lower in bothPHF6: predicted G418 readthrough 0.4%; measured change after the stop +0.97 percentage points; Lower in bothHS6ST2: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothFPGT: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothRRP12: predicted G418 readthrough 0.7%; measured change after the stop +0.77 percentage points; Lower in bothFRAT2: predicted G418 readthrough 2.8%; measured change after the stop +9.73 percentage points; High in model + measurementGNAS: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothJUN: predicted G418 readthrough 0.8%; measured change after the stop +0.12 percentage points; High in model onlyALDH18A1: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothDHCR24: predicted G418 readthrough 0.5%; measured change after the stop +0.76 percentage points; Lower in bothMCTS1: predicted G418 readthrough 0.7%; measured change after the stop −0.39 percentage points; Lower in bothCUL4B: predicted G418 readthrough 0.5%; measured change after the stop −0.74 percentage points; Lower in bothNKAP: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothNDUFA1: predicted G418 readthrough 0.4%; measured change after the stop +1.11 percentage points; Lower in bothRNF113A: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothMRPL41: predicted G418 readthrough 1.1%; measured change after the stop +0.00 percentage points; High in model onlyMAGOH: predicted G418 readthrough 0.4%; measured change after the stop +0.68 percentage points; Lower in bothCDC5L: predicted G418 readthrough 0.9%; measured change after the stop +0.82 percentage points; High in model onlyCEP55: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; High in model onlyCOA7: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothTNKS2: predicted G418 readthrough 0.5%; measured change after the stop +0.31 percentage points; Lower in bothHSP90AB1: predicted G418 readthrough 0.6%; measured change after the stop +0.17 percentage points; Lower in bothKIF20B: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothIFIT5: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothAMOT: predicted G418 readthrough 0.3%; measured change after the stop +0.17 percentage points; Lower in bothSURF4: predicted G418 readthrough 0.6%; measured change after the stop +3.03 percentage points; Lower in bothTP53RK: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothIRS4: predicted G418 readthrough 1.0%; measured change after the stop +0.61 percentage points; High in model onlyGHITM: predicted G418 readthrough 0.7%; measured change after the stop +0.84 percentage points; Lower in bothGTF3C4: predicted G418 readthrough 0.5%; measured change after the stop +0.39 percentage points; Lower in bothMAD2L1BP: predicted G418 readthrough 0.5%; measured change after the stop −1.44 percentage points; Lower in bothKIF2C: predicted G418 readthrough 1.4%; measured change after the stop +0.00 percentage points; High in model onlyRPS24: predicted G418 readthrough 0.5%; measured change after the stop +1.21 percentage points; Lower in bothZSWIM1: predicted G418 readthrough 2.0%; measured change after the stop +2.60 percentage points; High in model onlyZNF503: predicted G418 readthrough 0.5%; measured change after the stop +0.62 percentage points; Lower in bothRADX: predicted G418 readthrough 0.4%; measured change after the stop +0.03 percentage points; Lower in bothNUP188: predicted G418 readthrough 0.4%; measured change after the stop +0.52 percentage points; Lower in bothDOLK: predicted G418 readthrough 1.6%; measured change after the stop +1.40 percentage points; High in model onlyZMPSTE24: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothCHCHD1: predicted G418 readthrough 1.3%; measured change after the stop +1.68 percentage points; High in model onlyTBCC: predicted G418 readthrough 0.4%; measured change after the stop −0.11 percentage points; Lower in bothTRUB2: predicted G418 readthrough 0.6%; measured change after the stop +1.41 percentage points; Lower in bothMRPS16: predicted G418 readthrough 0.3%; measured change after the stop −0.43 percentage points; Lower in bothNDUFS5: predicted G418 readthrough 0.7%; measured change after the stop +1.64 percentage points; Lower in bothUTP11: predicted G418 readthrough 2.4%; measured change after the stop +6.18 percentage points; High in model + measurementSF3A3: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothTSPAN6: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothYRDC: predicted G418 readthrough 0.8%; measured change after the stop +4.08 percentage points; High in model onlyCDCA8: predicted G418 readthrough 0.7%; measured change after the stop −0.50 percentage points; Lower in bothNAP1L3: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothPSMB2: predicted G418 readthrough 0.9%; measured change after the stop +0.67 percentage points; High in model onlyNCDN: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothCDK9: predicted G418 readthrough 0.9%; measured change after the stop +5.01 percentage points; High in model + measurementBRWD3: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothMAFB: predicted G418 readthrough 0.6%; measured change after the stop +3.30 percentage points; Lower in bothPGK1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothATRX: predicted G418 readthrough 0.9%; measured change after the stop +0.00 percentage points; High in model onlyGLO1: predicted G418 readthrough 0.5%; measured change after the stop +2.29 percentage points; Lower in bothRBBP4: predicted G418 readthrough 0.5%; measured change after the stop +0.62 percentage points; Lower in bothPPP6C: predicted G418 readthrough 1.1%; measured change after the stop +0.70 percentage points; High in model onlyHDAC1: predicted G418 readthrough 1.1%; measured change after the stop +3.33 percentage points; High in model onlyKPNA6: predicted G418 readthrough 0.7%; measured change after the stop +0.68 percentage points; Lower in bothPIN4: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothPPIL1: predicted G418 readthrough 0.3%; measured change after the stop +0.63 percentage points; Lower in bothPEF1: predicted G418 readthrough 3.3%; measured change after the stop +6.14 percentage points; High in model + measurementSRSF3: predicted G418 readthrough 0.5%; measured change after the stop +3.48 percentage points; Lower in bothNDUFA8: predicted G418 readthrough 0.3%; measured change after the stop +1.27 percentage points; Lower in bothYTHDF2: predicted G418 readthrough 0.3%; measured change after the stop +0.45 percentage points; Lower in bothSRPK1: predicted G418 readthrough 0.4%; measured change after the stop +0.89 percentage points; Lower in bothRPA2: predicted G418 readthrough 0.6%; measured change after the stop −0.72 percentage points; Lower in bothATP6V1G1: predicted G418 readthrough 0.6%; measured change after the stop +1.96 percentage points; Lower in bothROMO1: predicted G418 readthrough 0.6%; measured change after the stop +1.16 percentage points; Lower in bothRBM12: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothPOLE3: predicted G418 readthrough 0.8%; measured change after the stop +1.32 percentage points; Lower in bothSLC31A1: predicted G418 readthrough 3.2%; measured change after the stop +1.26 percentage points; High in model onlySELENON: predicted G418 readthrough 0.4%; measured change after the stop +1.61 percentage points; Lower in bothCLIC4: predicted G418 readthrough 0.3%; measured change after the stop +1.72 percentage points; Lower in bothTXN: predicted G418 readthrough 0.3%; measured change after the stop +0.14 percentage points; Lower in bothID3: predicted G418 readthrough 1.7%; measured change after the stop +33.32 percentage points; High in model + measurementPJA1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothPFDN6: predicted G418 readthrough 3.4%; measured change after the stop +0.00 percentage points; High in model onlyWDR46: predicted G418 readthrough 0.7%; measured change after the stop +0.63 percentage points; Lower in bothFZD8: predicted G418 readthrough 0.5%; measured change after the stop +1.48 percentage points; Lower in bothSMC2: predicted G418 readthrough 0.3%; measured change after the stop +0.66 percentage points; Lower in bothBRD2: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothMRPL50: predicted G418 readthrough 0.3%; measured change after the stop +2.99 percentage points; Lower in bothAMER1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothSPIN4: predicted G418 readthrough 0.3%; measured change after the stop +0.31 percentage points; Lower in bothZXDB: predicted G418 readthrough 0.9%; measured change after the stop +2.12 percentage points; High in model onlyNCBP1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothUBR4: predicted G418 readthrough 1.9%; measured change after the stop +0.36 percentage points; High in model onlySDHB: predicted G418 readthrough 0.6%; measured change after the stop +0.63 percentage points; Lower in bothZBTB12: predicted G418 readthrough 0.7%; measured change after the stop +1.47 percentage points; Lower in bothSDHD: predicted G418 readthrough 2.5%; measured change after the stop +5.00 percentage points; High in model + measurementHSPA1B: predicted G418 readthrough 0.7%; measured change after the stop +3.15 percentage points; Lower in bothHSPA1A: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothHSPA1L: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothVARS1: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyDYNC2H1: predicted G418 readthrough 0.3%; measured change after the stop +0.58 percentage points; Lower in bothSPEN: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothCLIC1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothCSNK2B: predicted G418 readthrough 1.1%; measured change after the stop +1.78 percentage points; High in model onlyPRRC2A: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothHLA-C: predicted G418 readthrough 1.0%; measured change after the stop +1.95 percentage points; High in model onlyHNRNPK: predicted G418 readthrough 0.6%; measured change after the stop +2.45 percentage points; Lower in bothZIC2: predicted G418 readthrough 1.2%; measured change after the stop +3.09 percentage points; High in model onlyTM9SF2: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothUBQLN1: predicted G418 readthrough 1.0%; measured change after the stop +0.11 percentage points; High in model onlyMDC1: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothOTUD1: predicted G418 readthrough 1.1%; measured change after the stop +7.08 percentage points; High in model + measurementPIM2: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothPSAT1: predicted G418 readthrough 1.0%; measured change after the stop +0.76 percentage points; High in model onlyPPP1CA: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyRBM3: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothUBIAD1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothC9orf40: predicted G418 readthrough 1.4%; measured change after the stop +2.32 percentage points; High in model onlyCST3: predicted G418 readthrough 0.7%; measured change after the stop +3.26 percentage points; Lower in bothSOX21: predicted G418 readthrough 3.3%; measured change after the stop +9.98 percentage points; High in model + measurementSRM: predicted G418 readthrough 1.1%; measured change after the stop +0.00 percentage points; High in model onlyGPC6: predicted G418 readthrough 0.2%; measured change after the stop +2.00 percentage points; Lower in bothTRIM27: predicted G418 readthrough 0.5%; measured change after the stop +1.83 percentage points; Lower in bothPOU4F1: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothARL5B: predicted G418 readthrough 0.3%; measured change after the stop +0.97 percentage points; Lower in bothSF1: predicted G418 readthrough 0.4%; measured change after the stop +1.18 percentage points; Lower in bothKCTD12: predicted G418 readthrough 0.4%; measured change after the stop +4.52 percentage points; Lower in bothUCHL3: predicted G418 readthrough 1.0%; measured change after the stop −0.62 percentage points; High in model onlyTBC1D4: predicted G418 readthrough 0.2%; measured change after the stop +0.62 percentage points; Lower in bothALDH1B1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothNOL9: predicted G418 readthrough 1.2%; measured change after the stop +0.00 percentage points; High in model onlyH4C8: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothH2BC10: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothDIS3: predicted G418 readthrough 1.3%; measured change after the stop +0.00 percentage points; High in model onlyH2BC5: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothH2AC6: predicted G418 readthrough 0.9%; measured change after the stop +0.00 percentage points; High in model onlyH4C3: predicted G418 readthrough 0.6%; measured change after the stop +1.22 percentage points; Lower in bothNDUFB11: predicted G418 readthrough 1.5%; measured change after the stop +2.26 percentage points; High in model onlyRRBP1: predicted G418 readthrough 0.6%; measured change after the stop +0.81 percentage points; Lower in bothMZT1: predicted G418 readthrough 2.3%; measured change after the stop +21.84 percentage points; High in model + measurementRTN3: predicted G418 readthrough 0.3%; measured change after the stop +0.86 percentage points; Lower in bothPCDH9: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothAHNAK: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothCKAP4: predicted G418 readthrough 0.4%; measured change after the stop +0.33 percentage points; Lower in bothUSP9X: predicted G418 readthrough 0.9%; measured change after the stop +0.00 percentage points; High in model onlyAPRT: predicted G418 readthrough 1.5%; measured change after the stop +0.00 percentage points; High in model onlyBCOR: predicted G418 readthrough 0.5%; measured change after the stop +0.79 percentage points; Lower in bothSKI: predicted G418 readthrough 0.3%; measured change after the stop +1.00 percentage points; Lower in bothUQCRQ: predicted G418 readthrough 0.6%; measured change after the stop +2.03 percentage points; Lower in bothID4: predicted G418 readthrough 0.7%; measured change after the stop +3.92 percentage points; Lower in bothMCM10: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothESD: predicted G418 readthrough 1.0%; measured change after the stop +0.95 percentage points; High in model onlyZFC3H1: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothSDHAF1: predicted G418 readthrough 1.8%; measured change after the stop +5.80 percentage points; High in model + measurementRPP25L: predicted G418 readthrough 0.5%; measured change after the stop +12.70 percentage points; High in measurement onlyPCNA: predicted G418 readthrough 0.5%; measured change after the stop +1.41 percentage points; Lower in bothB3GALT6: predicted G418 readthrough 0.6%; measured change after the stop +6.25 percentage points; High in measurement onlyPRDX4: predicted G418 readthrough 1.0%; measured change after the stop +0.34 percentage points; High in model onlyKBTBD7: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothKBTBD6: predicted G418 readthrough 0.6%; measured change after the stop +0.81 percentage points; Lower in bothEIF1AX: predicted G418 readthrough 0.3%; measured change after the stop +1.33 percentage points; Lower in bothREEP5: predicted G418 readthrough 0.3%; measured change after the stop −1.46 percentage points; Lower in bothIPO7: predicted G418 readthrough 1.0%; measured change after the stop +0.67 percentage points; High in model onlyCENPB: predicted G418 readthrough 1.1%; measured change after the stop +1.76 percentage points; High in model onlyTXNDC5: predicted G418 readthrough 0.7%; measured change after the stop +1.15 percentage points; Lower in bothNDUFB6: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothRFC3: predicted G418 readthrough 2.0%; measured change after the stop +2.86 percentage points; High in model onlyRBBP7: predicted G418 readthrough 0.4%; measured change after the stop +1.61 percentage points; Lower in bothSYAP1: predicted G418 readthrough 1.5%; measured change after the stop +1.79 percentage points; High in model onlyGDI2: predicted G418 readthrough 0.9%; measured change after the stop +0.00 percentage points; High in model onlyFASTKD5: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyMRPS26: predicted G418 readthrough 0.7%; measured change after the stop +1.14 percentage points; Lower in bothTBCA: predicted G418 readthrough 1.3%; measured change after the stop +2.02 percentage points; High in model onlyRPS6: predicted G418 readthrough 0.3%; measured change after the stop −0.21 percentage points; Lower in bothZNF768: predicted G418 readthrough 1.7%; measured change after the stop +0.44 percentage points; High in model onlyHAUS6: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothRRAGA: predicted G418 readthrough 0.9%; measured change after the stop +3.94 percentage points; High in model onlyBTF3: predicted G418 readthrough 0.6%; measured change after the stop +1.26 percentage points; Lower in bothSLC7A1: predicted G418 readthrough 1.6%; measured change after the stop +7.33 percentage points; High in model + measurementEIF3CL: predicted G418 readthrough 0.5%; measured change after the stop +0.64 percentage points; Lower in bothSELENOO: predicted G418 readthrough 0.5%; measured change after the stop +0.61 percentage points; Lower in bothITPR2: predicted G418 readthrough 1.6%; measured change after the stop +0.00 percentage points; High in model onlySNRPB: predicted G418 readthrough 3.6%; measured change after the stop +0.00 percentage points; High in model onlyIDI1: predicted G418 readthrough 0.7%; measured change after the stop −0.54 percentage points; Lower in bothSLC25A6: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothCOMMD8: predicted G418 readthrough 0.6%; measured change after the stop +0.05 percentage points; Lower in bothZZEF1: predicted G418 readthrough 1.3%; measured change after the stop +0.00 percentage points; High in model onlyGART: predicted G418 readthrough 0.7%; measured change after the stop +1.92 percentage points; Lower in bothPLPP6: predicted G418 readthrough 2.2%; measured change after the stop +1.07 percentage points; High in model onlyPGM2: predicted G418 readthrough 0.4%; measured change after the stop +1.00 percentage points; Lower in bothSACS: predicted G418 readthrough 0.4%; measured change after the stop +1.40 percentage points; Lower in bothFSCN1: predicted G418 readthrough 0.5%; measured change after the stop +1.11 percentage points; Lower in bothSAP18: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothMRFAP1: predicted G418 readthrough 0.5%; measured change after the stop +3.67 percentage points; Lower in bothURB1: predicted G418 readthrough 2.4%; measured change after the stop +0.00 percentage points; High in model onlyCOPS2: predicted G418 readthrough 0.9%; measured change after the stop +0.00 percentage points; High in model onlyHSPD1: predicted G418 readthrough 0.7%; measured change after the stop +0.19 percentage points; Lower in bothNCBP3: predicted G418 readthrough 1.9%; measured change after the stop +0.00 percentage points; High in model onlyDIAPH1: predicted G418 readthrough 0.6%; measured change after the stop +0.79 percentage points; Lower in bothRNF20: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyGARS1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothEP400: predicted G418 readthrough 0.3%; measured change after the stop +0.70 percentage points; Lower in bothCRYBG3: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothRTF1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothTSN: predicted G418 readthrough 0.5%; measured change after the stop +1.63 percentage points; Lower in bothRNF216: predicted G418 readthrough 1.9%; measured change after the stop +0.00 percentage points; High in model onlyEID2: predicted G418 readthrough 0.5%; measured change after the stop +2.29 percentage points; Lower in bothRTL8B: predicted G418 readthrough 0.7%; measured change after the stop +7.66 percentage points; High in measurement onlyPTOV1: predicted G418 readthrough 0.2%; measured change after the stop +0.43 percentage points; Lower in bothRPL13A: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothHNRNPUL1: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothXRCC5: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothFAM117B: predicted G418 readthrough 0.6%; measured change after the stop +3.10 percentage points; Lower in bothMB21D2: predicted G418 readthrough 1.1%; measured change after the stop +0.00 percentage points; High in model onlyRPLP0: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothHNRNPA3: predicted G418 readthrough 0.5%; measured change after the stop +1.94 percentage points; Lower in bothHOXD13: predicted G418 readthrough 1.4%; measured change after the stop +2.56 percentage points; High in model onlyKLHL23: predicted G418 readthrough 0.3%; measured change after the stop +0.81 percentage points; Lower in bothECT2: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothSTT3A: predicted G418 readthrough 0.3%; measured change after the stop +0.78 percentage points; Lower in bothTDG: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothMTPN: predicted G418 readthrough 1.0%; measured change after the stop +4.84 percentage points; High in model onlyNME1: predicted G418 readthrough 2.2%; measured change after the stop +13.89 percentage points; High in model + measurementPTGFRN: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothPLXNA1: predicted G418 readthrough 1.1%; measured change after the stop +0.39 percentage points; High in model onlyHNRNPH1: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothCOASY: predicted G418 readthrough 1.1%; measured change after the stop +3.25 percentage points; High in model onlyCNP: predicted G418 readthrough 0.5%; measured change after the stop +0.52 percentage points; Lower in bothMRPL19: predicted G418 readthrough 1.5%; measured change after the stop +2.96 percentage points; High in model onlyHMMR: predicted G418 readthrough 0.3%; measured change after the stop +0.77 percentage points; Lower in bothPABIR1: predicted G418 readthrough 0.9%; measured change after the stop +2.85 percentage points; High in model onlyATP5MC2: predicted G418 readthrough 0.6%; measured change after the stop +0.83 percentage points; Lower in bothCCT4: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothRPL34: predicted G418 readthrough 0.4%; measured change after the stop +0.54 percentage points; Lower in bothZNF207: predicted G418 readthrough 1.0%; measured change after the stop +0.55 percentage points; High in model onlyMATR3: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothINCENP: predicted G418 readthrough 0.7%; measured change after the stop +1.66 percentage points; Lower in bothRCN2: predicted G418 readthrough 0.6%; measured change after the stop +0.60 percentage points; Lower in bothCDC23: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothKIF20A: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothPSAP: predicted G418 readthrough 0.4%; measured change after the stop +1.55 percentage points; Lower in bothC15orf39: predicted G418 readthrough 0.8%; measured change after the stop −0.45 percentage points; Lower in bothDNAJC30: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothCDK1: predicted G418 readthrough 0.7%; measured change after the stop +1.43 percentage points; Lower in bothMAPK1IP1L: predicted G418 readthrough 0.4%; measured change after the stop +0.91 percentage points; Lower in bothPURB: predicted G418 readthrough 1.3%; measured change after the stop +9.59 percentage points; High in model + measurementYWHAZ: predicted G418 readthrough 0.7%; measured change after the stop +0.87 percentage points; Lower in bothFKBP3: predicted G418 readthrough 1.2%; measured change after the stop +2.60 percentage points; High in model onlyDDX39B: predicted G418 readthrough 0.4%; measured change after the stop +0.85 percentage points; Lower in bothRPL14: predicted G418 readthrough 0.4%; measured change after the stop +0.56 percentage points; Lower in bothID2: predicted G418 readthrough 0.7%; measured change after the stop +2.93 percentage points; Lower in bothCBX3: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothPSMA1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothRPS8: predicted G418 readthrough 0.3%; measured change after the stop +0.20 percentage points; Lower in bothTPI1: predicted G418 readthrough 0.7%; measured change after the stop +1.57 percentage points; Lower in bothH2BC4: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothNUDT19: predicted G418 readthrough 0.3%; measured change after the stop −0.41 percentage points; Lower in bothCCNF: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothPRKAA1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothSMU1: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothMRPL23: predicted G418 readthrough 1.7%; measured change after the stop +2.85 percentage points; High in model onlyMRPS34: predicted G418 readthrough 0.4%; measured change after the stop +0.33 percentage points; Lower in bothFAM98B: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothALG3: predicted G418 readthrough 0.6%; measured change after the stop −0.71 percentage points; Lower in bothMED13: predicted G418 readthrough 0.9%; measured change after the stop +0.61 percentage points; High in model onlyTTC28: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothDDT: predicted G418 readthrough 1.8%; measured change after the stop +8.96 percentage points; High in model + measurementUSP34: predicted G418 readthrough 2.7%; measured change after the stop +0.00 percentage points; High in model onlyGSTP1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothDOT1L: predicted G418 readthrough 0.5%; measured change after the stop +1.56 percentage points; Lower in bothATP5F1A: predicted G418 readthrough 0.5%; measured change after the stop +0.43 percentage points; Lower in bothSEC24A: predicted G418 readthrough 0.8%; measured change after the stop +1.96 percentage points; Lower in bothATL3: predicted G418 readthrough 1.2%; measured change after the stop +4.29 percentage points; High in model onlyTOMM6: predicted G418 readthrough 0.5%; measured change after the stop +3.81 percentage points; Lower in bothNOMO3: predicted G418 readthrough 0.4%; measured change after the stop +1.82 percentage points; Lower in bothKNL1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothKDM5A: predicted G418 readthrough 2.1%; measured change after the stop +0.81 percentage points; High in model onlyROCK1: predicted G418 readthrough 1.3%; measured change after the stop +0.00 percentage points; High in model onlyFKBP1A: predicted G418 readthrough 2.2%; measured change after the stop +10.74 percentage points; High in model + measurementMMACHC: predicted G418 readthrough 1.0%; measured change after the stop +0.75 percentage points; High in model onlySNU13: predicted G418 readthrough 0.3%; measured change after the stop +0.73 percentage points; Lower in bothAKAP12: predicted G418 readthrough 0.5%; measured change after the stop +0.21 percentage points; Lower in bothCPE: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothDDX17: predicted G418 readthrough 0.9%; measured change after the stop +0.41 percentage points; High in model onlyIMP3: predicted G418 readthrough 1.3%; measured change after the stop +4.40 percentage points; High in model onlyH3C15: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothPSME4: predicted G418 readthrough 0.5%; measured change after the stop +0.94 percentage points; Lower in bothINTS1: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothSMS: predicted G418 readthrough 0.5%; measured change after the stop +3.01 percentage points; Lower in bothPRAME: predicted G418 readthrough 1.5%; measured change after the stop +2.47 percentage points; High in model onlySMCR8: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothRPS14: predicted G418 readthrough 0.6%; measured change after the stop +0.04 percentage points; Lower in bothCBX6: predicted G418 readthrough 0.5%; measured change after the stop +2.16 percentage points; Lower in bothWDR43: predicted G418 readthrough 0.6%; measured change after the stop +4.09 percentage points; Lower in bothELAVL1: predicted G418 readthrough 0.5%; measured change after the stop +0.83 percentage points; Lower in bothPTMA: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothBZW1: predicted G418 readthrough 0.9%; measured change after the stop +0.89 percentage points; High in model onlyPOTEJ: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothJPT1: predicted G418 readthrough 0.9%; measured change after the stop +1.34 percentage points; High in model onlyRAB1A: predicted G418 readthrough 0.3%; measured change after the stop −0.04 percentage points; Lower in bothELOB: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothPOTEF: predicted G418 readthrough 0.4%; measured change after the stop −0.47 percentage points; Lower in bothDAZAP2: predicted G418 readthrough 0.4%; measured change after the stop +3.00 percentage points; Lower in bothWDR26: predicted G418 readthrough 2.8%; measured change after the stop +0.00 percentage points; High in model onlySNRPE: predicted G418 readthrough 0.4%; measured change after the stop +0.64 percentage points; Lower in bothNOP14: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothKBTBD8: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothIK: predicted G418 readthrough 0.3%; measured change after the stop +0.94 percentage points; Lower in bothUSP19: predicted G418 readthrough 1.4%; measured change after the stop +0.00 percentage points; High in model onlyRHOA: predicted G418 readthrough 0.8%; measured change after the stop +5.70 percentage points; High in measurement onlyGNL3: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothPRR12: predicted G418 readthrough 1.3%; measured change after the stop +0.00 percentage points; High in model onlyGPX1: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothSTRAP: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothTSPYL4: predicted G418 readthrough 0.3%; measured change after the stop +2.65 percentage points; Lower in bothSUMO2: predicted G418 readthrough 0.8%; measured change after the stop +1.75 percentage points; Lower in bothPERP: predicted G418 readthrough 0.6%; measured change after the stop +3.31 percentage points; Lower in bothBIRC6: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothZNF697: predicted G418 readthrough 0.7%; measured change after the stop +1.96 percentage points; Lower in bothCMSS1: predicted G418 readthrough 0.4%; measured change after the stop +0.87 percentage points; Lower in bothPDHA1: predicted G418 readthrough 0.8%; measured change after the stop +1.63 percentage points; Lower in bothCIAO2B: predicted G418 readthrough 0.9%; measured change after the stop +6.35 percentage points; High in model + measurementLDHA: predicted G418 readthrough 0.4%; measured change after the stop +0.62 percentage points; Lower in bothRPL23A: predicted G418 readthrough 0.3%; measured change after the stop +0.06 percentage points; Lower in bothGLG1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothIRGQ: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothTOP2A: predicted G418 readthrough 0.5%; measured change after the stop +0.21 percentage points; Lower in bothCOPG2: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothMORF4L1: predicted G418 readthrough 0.5%; measured change after the stop +1.55 percentage points; Lower in bothTMEM185B: predicted G418 readthrough 0.5%; measured change after the stop +4.26 percentage points; Lower in bothEFTUD2: predicted G418 readthrough 0.7%; measured change after the stop +0.88 percentage points; Lower in bothTOMM40: predicted G418 readthrough 0.5%; measured change after the stop +2.15 percentage points; Lower in bothFTSJ3: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothRPL32: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothRANBP1: predicted G418 readthrough 0.3%; measured change after the stop +0.23 percentage points; Lower in bothFAM136A: predicted G418 readthrough 0.3%; measured change after the stop +2.19 percentage points; Lower in bothTNRC18: predicted G418 readthrough 0.7%; measured change after the stop +0.54 percentage points; Lower in bothSUPT5H: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyPCYOX1: predicted G418 readthrough 0.9%; measured change after the stop +1.32 percentage points; High in model onlyHSBP1: predicted G418 readthrough 1.5%; measured change after the stop +20.70 percentage points; High in model + measurementPOM121: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothGSPT1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothEXOSC6: predicted G418 readthrough 0.7%; measured change after the stop +2.43 percentage points; Lower in bothDNTTIP2: predicted G418 readthrough 0.4%; measured change after the stop +0.48 percentage points; Lower in bothTMA7: predicted G418 readthrough 0.2%; measured change after the stop −0.00 percentage points; Lower in bothDHFR: predicted G418 readthrough 0.7%; measured change after the stop −0.31 percentage points; Lower in bothCOPB1: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothFAT1: predicted G418 readthrough 1.6%; measured change after the stop +0.72 percentage points; High in model onlyUQCC4: predicted G418 readthrough 1.8%; measured change after the stop +45.78 percentage points; High in model + measurementUBA52: predicted G418 readthrough 0.2%; measured change after the stop +1.04 percentage points; Lower in bothHERC1: predicted G418 readthrough 1.9%; measured change after the stop +0.00 percentage points; High in model onlyRIF1: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothSMG1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothCDK12: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothDCTN4: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothEIF4E: predicted G418 readthrough 0.3%; measured change after the stop +0.58 percentage points; Lower in bothNOL7: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothPOTEI: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothDDX46: predicted G418 readthrough 0.5%; measured change after the stop +0.46 percentage points; Lower in bothTRA2B: predicted G418 readthrough 0.4%; measured change after the stop +1.18 percentage points; Lower in bothGTSE1: predicted G418 readthrough 0.4%; measured change after the stop +0.38 percentage points; Lower in bothPRMT1: predicted G418 readthrough 0.7%; measured change after the stop +1.65 percentage points; Lower in bothRFC5: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothNACA: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothZBTB10: predicted G418 readthrough 0.8%; measured change after the stop +0.90 percentage points; High in model onlySYPL1: predicted G418 readthrough 0.3%; measured change after the stop +1.00 percentage points; Lower in bothSTMN1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothTRRAP: predicted G418 readthrough 1.5%; measured change after the stop +0.00 percentage points; High in model onlyTMX1: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothPGAM4: predicted G418 readthrough 0.6%; measured change after the stop +1.19 percentage points; Lower in bothBAP1: predicted G418 readthrough 0.4%; measured change after the stop +1.78 percentage points; Lower in bothTKT: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothROBO1: predicted G418 readthrough 0.9%; measured change after the stop +0.00 percentage points; High in model onlyNAA10: predicted G418 readthrough 0.6%; measured change after the stop −0.50 percentage points; Lower in bothPCMT1: predicted G418 readthrough 0.9%; measured change after the stop +2.86 percentage points; High in model onlyANP32A: predicted G418 readthrough 0.4%; measured change after the stop +1.57 percentage points; Lower in bothPPIA: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothEIF1: predicted G418 readthrough 0.5%; measured change after the stop +1.58 percentage points; Lower in bothSELENOT: predicted G418 readthrough 0.9%; measured change after the stop +1.71 percentage points; High in model onlyATP6V0B: predicted G418 readthrough 0.5%; measured change after the stop +4.18 percentage points; Lower in bothU2SURP: predicted G418 readthrough 2.2%; measured change after the stop +3.15 percentage points; High in model onlyPTPRG: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyMRPS14: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothFAM162A: predicted G418 readthrough 0.7%; measured change after the stop +1.59 percentage points; Lower in bothSEPHS2: predicted G418 readthrough 0.6%; measured change after the stop +1.96 percentage points; Lower in bothRPL23: predicted G418 readthrough 1.4%; measured change after the stop +1.67 percentage points; High in model onlyTCAF1: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothCHCHD10: predicted G418 readthrough 0.5%; measured change after the stop +3.36 percentage points; Lower in bothRPSA2: predicted G418 readthrough 0.4%; measured change after the stop −0.39 percentage points; Lower in bothTFAM: predicted G418 readthrough 0.4%; measured change after the stop +0.60 percentage points; Lower in bothCIAO1: predicted G418 readthrough 0.8%; measured change after the stop +0.56 percentage points; Lower in bothSLBP: predicted G418 readthrough 0.9%; measured change after the stop +2.92 percentage points; High in model onlyRPL37A: predicted G418 readthrough 0.3%; measured change after the stop +1.27 percentage points; Lower in bothNUDT5: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothRPL7L1: predicted G418 readthrough 0.5%; measured change after the stop +1.31 percentage points; Lower in bothEBP: predicted G418 readthrough 0.5%; measured change after the stop +6.20 percentage points; High in measurement onlySELENOK: predicted G418 readthrough 0.4%; measured change after the stop +12.19 percentage points; High in measurement onlyDHX36: predicted G418 readthrough 3.5%; measured change after the stop −0.04 percentage points; High in model onlyNDUFA6: predicted G418 readthrough 0.5%; measured change after the stop +1.79 percentage points; Lower in bothPGAM5: predicted G418 readthrough 0.8%; measured change after the stop +2.99 percentage points; Lower in bothZCCHC3: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothSPCS3: predicted G418 readthrough 0.4%; measured change after the stop +2.99 percentage points; Lower in bothANKRD50: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothTIMM8B: predicted G418 readthrough 0.4%; measured change after the stop −0.25 percentage points; Lower in bothSLIT2: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothTIGD5: predicted G418 readthrough 1.4%; measured change after the stop +10.64 percentage points; High in model + measurementTMEM33: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothALYREF: predicted G418 readthrough 0.7%; measured change after the stop +1.32 percentage points; Lower in bothLUC7L3: predicted G418 readthrough 0.5%; measured change after the stop +0.74 percentage points; Lower in bothMRPS30: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothNDUFA13: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothPAICS: predicted G418 readthrough 0.5%; measured change after the stop +0.63 percentage points; Lower in bothNME2: predicted G418 readthrough 0.3%; measured change after the stop +0.51 percentage points; Lower in bothRACK1: predicted G418 readthrough 0.5%; measured change after the stop +0.11 percentage points; Lower in bothWDR36: predicted G418 readthrough 0.3%; measured change after the stop +1.17 percentage points; Lower in bothUSO1: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothCPNE3: predicted G418 readthrough 1.6%; measured change after the stop +3.18 percentage points; High in model onlyLARP1: predicted G418 readthrough 0.7%; measured change after the stop +1.14 percentage points; Lower in bothHDAC2: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothATP6V0E1: predicted G418 readthrough 0.5%; measured change after the stop +7.72 percentage points; High in measurement onlyATXN7L3B: predicted G418 readthrough 0.8%; measured change after the stop +5.49 percentage points; High in model + measurementUBR5: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothEIF5AL1: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothLAPTM4B: predicted G418 readthrough 0.4%; measured change after the stop −0.26 percentage points; Lower in bothNACA2: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothEIF3H: predicted G418 readthrough 0.3%; measured change after the stop +1.30 percentage points; Lower in bothDPYSL2: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothINTS8: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothTXNRD1: predicted G418 readthrough 0.3%; measured change after the stop +7.18 percentage points; High in measurement onlySELENOS: predicted G418 readthrough 0.2%; measured change after the stop +0.00 percentage points; Lower in bothCHMP1B: predicted G418 readthrough 1.2%; measured change after the stop +0.13 percentage points; High in model onlyMANF: predicted G418 readthrough 0.3%; measured change after the stop +3.05 percentage points; Lower in bothCKAP5: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothFAU: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothEID1: predicted G418 readthrough 0.4%; measured change after the stop +1.40 percentage points; Lower in bothH2AX: predicted G418 readthrough 0.3%; measured change after the stop +0.99 percentage points; Lower in bothTPT1: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyBRK1: predicted G418 readthrough 0.4%; measured change after the stop +9.81 percentage points; High in measurement onlyEIF3M: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothRPS3: predicted G418 readthrough 0.8%; measured change after the stop +1.06 percentage points; Lower in bothAPI5: predicted G418 readthrough 0.8%; measured change after the stop +0.99 percentage points; Lower in bothAP5B1: predicted G418 readthrough 0.6%; measured change after the stop +0.53 percentage points; Lower in bothSELENOH: predicted G418 readthrough 0.4%; measured change after the stop +0.77 percentage points; Lower in bothNCAPD3: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothHSPA8: predicted G418 readthrough 0.4%; measured change after the stop +0.16 percentage points; Lower in bothPHB2: predicted G418 readthrough 0.7%; measured change after the stop +0.30 percentage points; Lower in bothSALL2: predicted G418 readthrough 0.5%; measured change after the stop +2.58 percentage points; Lower in bothEIF2S3B: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothOTUB1: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothATP2A2: predicted G418 readthrough 2.1%; measured change after the stop +0.36 percentage points; High in model onlyYTHDF3: predicted G418 readthrough 0.6%; measured change after the stop +0.23 percentage points; Lower in bothDMXL1: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyH4C16: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothBCORL1: predicted G418 readthrough 4.2%; measured change after the stop +0.26 percentage points; High in model onlyH2BC8: predicted G418 readthrough 0.3%; measured change after the stop +2.45 percentage points; Lower in bothPDCD2: predicted G418 readthrough 0.2%; measured change after the stop +0.90 percentage points; Lower in bothSLC1A5: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothRAN: predicted G418 readthrough 0.6%; measured change after the stop +0.29 percentage points; Lower in bothCHD4: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothVIM: predicted G418 readthrough 0.4%; measured change after the stop +0.20 percentage points; Lower in bothH2AJ: predicted G418 readthrough 2.4%; measured change after the stop +0.00 percentage points; High in model onlyCAND1: predicted G418 readthrough 0.9%; measured change after the stop +0.00 percentage points; High in model onlyDAPK3: predicted G418 readthrough 0.3%; measured change after the stop +0.93 percentage points; Lower in bothPRPF40A: predicted G418 readthrough 0.3%; measured change after the stop +0.47 percentage points; Lower in bothRPL41: predicted G418 readthrough 0.3%; measured change after the stop +15.03 percentage points; High in measurement onlyGATC: predicted G418 readthrough 1.2%; measured change after the stop −0.22 percentage points; High in model onlyNUP37: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothSLC25A3: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothRPL36A: predicted G418 readthrough 0.5%; measured change after the stop +0.29 percentage points; Lower in bothHNRNPC: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothGTF2A1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothTMPO: predicted G418 readthrough 0.7%; measured change after the stop +3.44 percentage points; Lower in bothMAP1LC3B2: predicted G418 readthrough 0.2%; measured change after the stop +0.55 percentage points; Lower in bothERH: predicted G418 readthrough 0.4%; measured change after the stop +0.89 percentage points; Lower in bothZBTB33: predicted G418 readthrough 0.3%; measured change after the stop +0.29 percentage points; Lower in bothTENT4B: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothRBM15B: predicted G418 readthrough 1.4%; measured change after the stop +0.88 percentage points; High in model onlyZNF106: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothRSL1D1: predicted G418 readthrough 0.3%; measured change after the stop +0.28 percentage points; Lower in bothACTG1: predicted G418 readthrough 0.4%; measured change after the stop +0.36 percentage points; Lower in bothH4C14: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothH4C15: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothRBM8A: predicted G418 readthrough 2.2%; measured change after the stop +9.43 percentage points; High in model + measurementANP32E: predicted G418 readthrough 0.6%; measured change after the stop +0.40 percentage points; Lower in bothUBL5: predicted G418 readthrough 0.5%; measured change after the stop +1.58 percentage points; Lower in bothSRSF8: predicted G418 readthrough 0.4%; measured change after the stop +4.66 percentage points; Lower in bothC19orf53: predicted G418 readthrough 0.7%; measured change after the stop +7.47 percentage points; High in measurement onlyILF3: predicted G418 readthrough 0.2%; measured change after the stop +0.60 percentage points; Lower in bothPSME3: predicted G418 readthrough 0.7%; measured change after the stop +2.42 percentage points; Lower in bothUQCR11: predicted G418 readthrough 0.8%; measured change after the stop +6.20 percentage points; High in measurement onlySPC24: predicted G418 readthrough 0.7%; measured change after the stop +4.32 percentage points; Lower in bothRNASEK: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothRUVBL2: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyTRAPPC5: predicted G418 readthrough 2.3%; measured change after the stop +0.45 percentage points; High in model onlyMCM3: predicted G418 readthrough 0.5%; measured change after the stop +0.54 percentage points; Lower in bothWDR83OS: predicted G418 readthrough 1.3%; measured change after the stop +1.65 percentage points; High in model onlyRPS19: predicted G418 readthrough 0.7%; measured change after the stop +0.47 percentage points; Lower in bothKHSRP: predicted G418 readthrough 1.3%; measured change after the stop +3.69 percentage points; High in model onlyRPS28: predicted G418 readthrough 0.5%; measured change after the stop +3.28 percentage points; Lower in bothLAMTOR5: predicted G418 readthrough 1.8%; measured change after the stop +8.93 percentage points; High in model + measurementEIF4G3: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothDDOST: predicted G418 readthrough 0.8%; measured change after the stop +1.02 percentage points; Lower in bothNCBP2AS2: predicted G418 readthrough 0.4%; measured change after the stop +4.19 percentage points; Lower in bothLIX1L: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothNOLC1: predicted G418 readthrough 1.2%; measured change after the stop +0.78 percentage points; High in model onlyH2AC19: predicted G418 readthrough 0.6%; measured change after the stop +0.73 percentage points; Lower in bothTIMM50: predicted G418 readthrough 0.8%; measured change after the stop +0.82 percentage points; Lower in bothWDR6: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothAP2B1: predicted G418 readthrough 0.7%; measured change after the stop +0.76 percentage points; Lower in bothNSA2: predicted G418 readthrough 2.3%; measured change after the stop +2.08 percentage points; High in model onlyZNF282: predicted G418 readthrough 0.8%; measured change after the stop +8.25 percentage points; High in measurement onlyNEFL: predicted G418 readthrough 1.1%; measured change after the stop +1.60 percentage points; High in model onlyLENG9: predicted G418 readthrough 1.6%; measured change after the stop +0.73 percentage points; High in model onlyMARCKS: predicted G418 readthrough 0.2%; measured change after the stop +0.12 percentage points; Lower in bothDCAF13: predicted G418 readthrough 0.4%; measured change after the stop +0.95 percentage points; Lower in bothH2BC15: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothH2AC16: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothDACH1: predicted G418 readthrough 0.7%; measured change after the stop +0.46 percentage points; Lower in bothELOA: predicted G418 readthrough 0.3%; measured change after the stop +0.75 percentage points; Lower in bothH3C1: predicted G418 readthrough 1.6%; measured change after the stop +1.52 percentage points; High in model onlyH2BC6: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothH4C4: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothCHD1: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothJADE3: predicted G418 readthrough 1.3%; measured change after the stop +1.54 percentage points; High in model onlyH4C5: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothH4C9: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothH2AC4: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothH2BC3: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothACACA: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothH3C11: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothRAVER1: predicted G418 readthrough 0.5%; measured change after the stop +0.00 percentage points; Lower in bothH4C1: predicted G418 readthrough 0.2%; measured change after the stop +0.20 percentage points; Lower in bothDDX52: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothH3C7: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothHNRNPA2B1: predicted G418 readthrough 0.5%; measured change after the stop +0.36 percentage points; Lower in bothMAGT1: predicted G418 readthrough 0.8%; measured change after the stop +0.13 percentage points; Lower in bothH4C13: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothNAP1L1: predicted G418 readthrough 0.5%; measured change after the stop +1.26 percentage points; Lower in bothH2AC15: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothPSMB3: predicted G418 readthrough 0.9%; measured change after the stop −0.60 percentage points; High in model onlyFAM83D: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothSPCS1: predicted G418 readthrough 0.8%; measured change after the stop +10.24 percentage points; High in measurement onlyVPS13D: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyH2BC14: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothH3C2: predicted G418 readthrough 0.5%; measured change after the stop +0.46 percentage points; Lower in bothMYC: predicted G418 readthrough 0.3%; measured change after the stop +0.48 percentage points; Lower in bothDDX24: predicted G418 readthrough 0.8%; measured change after the stop +0.43 percentage points; Lower in bothNOMO2: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothDCTN1: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothHNRNPU: predicted G418 readthrough 1.0%; measured change after the stop +0.23 percentage points; High in model onlyPHGDH: predicted G418 readthrough 0.7%; measured change after the stop +0.07 percentage points; Lower in bothPPT1: predicted G418 readthrough 1.2%; measured change after the stop +2.19 percentage points; High in model onlyALDOA: predicted G418 readthrough 0.3%; measured change after the stop +0.54 percentage points; Lower in bothRPL11: predicted G418 readthrough 0.3%; measured change after the stop −0.12 percentage points; Lower in bothSTEEP1: predicted G418 readthrough 0.6%; measured change after the stop +1.95 percentage points; Lower in bothDDX3X: predicted G418 readthrough 0.6%; measured change after the stop +1.67 percentage points; Lower in bothFOXF2: predicted G418 readthrough 0.6%; measured change after the stop +1.33 percentage points; Lower in bothSIX1: predicted G418 readthrough 0.4%; measured change after the stop +0.40 percentage points; Lower in bothFOXC1: predicted G418 readthrough 3.4%; measured change after the stop +9.89 percentage points; High in model + measurementFRRS1: predicted G418 readthrough 0.8%; measured change after the stop +2.96 percentage points; Lower in bothRPS26: predicted G418 readthrough 0.2%; measured change after the stop −0.13 percentage points; Lower in bothCHD8: predicted G418 readthrough 1.0%; measured change after the stop +0.00 percentage points; High in model onlyACTB: predicted G418 readthrough 0.5%; measured change after the stop +0.82 percentage points; Lower in bothRPL35A: predicted G418 readthrough 0.3%; measured change after the stop −0.14 percentage points; Lower in bothPRRC2C: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothITM2B: predicted G418 readthrough 0.9%; measured change after the stop +0.00 percentage points; High in model onlyRPS17: predicted G418 readthrough 0.7%; measured change after the stop +3.59 percentage points; Lower in bothJAGN1: predicted G418 readthrough 0.7%; measured change after the stop +1.71 percentage points; Lower in bothB2M: predicted G418 readthrough 0.3%; measured change after the stop +1.42 percentage points; Lower in bothRPS10: predicted G418 readthrough 0.2%; measured change after the stop +0.44 percentage points; Lower in bothHOXA13: predicted G418 readthrough 0.8%; measured change after the stop +3.61 percentage points; High in model onlyDARS2: predicted G418 readthrough 1.1%; measured change after the stop +3.70 percentage points; High in model onlyEIF4A3: predicted G418 readthrough 0.6%; measured change after the stop +0.20 percentage points; Lower in bothCOX6B1: predicted G418 readthrough 0.7%; measured change after the stop +4.39 percentage points; Lower in bothMCM4: predicted G418 readthrough 1.1%; measured change after the stop +0.14 percentage points; High in model onlyQSER1: predicted G418 readthrough 1.9%; measured change after the stop +0.24 percentage points; High in model onlyLRPAP1: predicted G418 readthrough 1.0%; measured change after the stop +2.56 percentage points; High in model onlyIGF1R: predicted G418 readthrough 1.0%; measured change after the stop +3.91 percentage points; High in model onlyCOX7B: predicted G418 readthrough 0.3%; measured change after the stop +0.22 percentage points; Lower in bothUHRF1: predicted G418 readthrough 0.8%; measured change after the stop +0.47 percentage points; High in model onlyCLUH: predicted G418 readthrough 0.5%; measured change after the stop +0.65 percentage points; Lower in bothZFHX4: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothEIF3F: predicted G418 readthrough 0.7%; measured change after the stop +1.28 percentage points; Lower in bothRPS27: predicted G418 readthrough 0.3%; measured change after the stop +0.39 percentage points; Lower in bothIPO5: predicted G418 readthrough 0.8%; measured change after the stop +0.22 percentage points; High in model onlyCITED1: predicted G418 readthrough 0.4%; measured change after the stop +2.09 percentage points; Lower in bothALX4: predicted G418 readthrough 1.8%; measured change after the stop +6.67 percentage points; High in model + measurementPDIA4: predicted G418 readthrough 1.1%; measured change after the stop +1.27 percentage points; High in model onlyMTHFD1: predicted G418 readthrough 0.4%; measured change after the stop −0.60 percentage points; Lower in bothRBMXL1: predicted G418 readthrough 0.4%; measured change after the stop +1.70 percentage points; Lower in bothDEK: predicted G418 readthrough 0.9%; measured change after the stop +0.57 percentage points; High in model onlyCDC42: predicted G418 readthrough 1.0%; measured change after the stop +5.04 percentage points; High in model + measurementAK2: predicted G418 readthrough 0.4%; measured change after the stop +0.74 percentage points; Lower in bothARHGAP35: predicted G418 readthrough 0.6%; measured change after the stop +1.20 percentage points; Lower in bothLDB1: predicted G418 readthrough 0.3%; measured change after the stop +0.60 percentage points; Lower in bothCELSR1: predicted G418 readthrough 0.7%; measured change after the stop +0.00 percentage points; Lower in bothPRKCSH: predicted G418 readthrough 1.6%; measured change after the stop +2.53 percentage points; High in model onlyEIF3I: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothSRPRB: predicted G418 readthrough 1.0%; measured change after the stop +3.18 percentage points; High in model onlyBRD4: predicted G418 readthrough 0.7%; measured change after the stop +1.61 percentage points; Lower in bothATP11C: predicted G418 readthrough 0.6%; measured change after the stop +2.43 percentage points; Lower in bothPOTEE: predicted G418 readthrough 0.7%; measured change after the stop +1.23 percentage points; Lower in bothPRRC2B: predicted G418 readthrough 3.1%; measured change after the stop +0.85 percentage points; High in model onlySLITRK5: predicted G418 readthrough 0.5%; measured change after the stop −0.74 percentage points; Lower in bothTASOR: predicted G418 readthrough 0.6%; measured change after the stop +0.00 percentage points; Lower in bothECPAS: predicted G418 readthrough 0.3%; measured change after the stop +0.00 percentage points; Lower in bothCOX7A2: predicted G418 readthrough 1.0%; measured change after the stop +0.92 percentage points; High in model onlySEC16A: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothH3C10: predicted G418 readthrough 1.0%; measured change after the stop +1.10 percentage points; High in model onlyH2AC25: predicted G418 readthrough 0.6%; measured change after the stop +0.47 percentage points; Lower in bothPCF11: predicted G418 readthrough 0.4%; measured change after the stop +0.00 percentage points; Lower in bothTASOR2: predicted G418 readthrough 1.5%; measured change after the stop +0.00 percentage points; High in model onlyGNG5B: predicted G418 readthrough 0.5%; measured change after the stop +0.93 percentage points; Lower in bothHNRNPA1L3: predicted G418 readthrough 0.8%; measured change after the stop +0.00 percentage points; Lower in bothMEASUREMENT HIGH ONLY · 40HIGH IN BOTH · 58 · MORE BURDENLOWER IN BOTH · 1,235 · NOT “SAFE”MODEL HIGH ONLY · 367MODEL “HIGH” CUT = TOP QUARTILE (0.8%)MEASUREMENT “HIGH” CUT = +5 PERCENTAGE POINTS8 VALUES ABOVE +15 POINTS ARE PINNED TO THE TOP EDGEmodel-predicted G418 readthrough (higher → more continuation)measured change after stop (percentage points)
  • high in model and measurement
  • high in model only
  • high in measurement only
  • lower in both

UP AND RIGHT = MORE UNINTENDED CONTINUATION · Lower-left means lower on these two measures, not proven safe.

Details & limits
“High” is descriptive: a measured increase above 5 percentage points in the share of ribosomes found after the stop, or the highest quarter of model predictions. These cuts were chosen to organise the atlas, not as biological or clinical safety thresholds. Hover a dot for the gene and exact values.
0.29rank correlation

1 means identical ranking; 0 means none. The 95% interval is 0.240.33.

03

Where the model disagrees

Four groups account for every stop. 407 cross the line.

Agreement means both values fall on the same side of descriptive cuts. Both-high still means greater burden.

Nearly one quarter are high on only one of the two measures

Complete partition of the same 1,700 matched normal stops

measured
04258501,2751,700AGREES · Both highDISAGREES · Model high onlyDISAGREES · Measurement high onlyAGREES · Lower in bothHigh in model + measurement: 58 of 1,700 (3.4%); model and measurement agree on the descriptive sideHigh in model only: 367 of 1,700 (21.6%); model and measurement disagreeHigh in measurement only: 40 of 1,700 (2.4%); model and measurement disagreeLower in both: 1,235 of 1,700 (72.6%); model and measurement agree on the descriptive side58 · 3.4%367 · 21.6%40 · 2.4%1,235 · 72.6%AGREEMENT SUPPORTS THE RANKINGDISAGREEMENT REQUIRES MEASUREMENTcanonical stops (same denominator: 1,700)
  • high in both · agreement, higher burden
  • model high only · disagreement
  • measurement high only · disagreement
  • lower in both · agreement, not a safety call

AGREEMENT SUPPORTS RANKING · Disagreement shows where direct measurement changes the model-only story.

Details & limits
The groups use the same post-analysis cuts as the scatter. They are a compact diagnostic, not a test of harm. In particular, “lower in both” does not demonstrate normal protein termination or lack of toxicity.
23.9%model–assay disagreement

407 stops change category when measurement replaces prediction.

04

Highest-value next experiment

G418 has an atlas. Every other route is an open question.

Only G418 has a measured atlas; five other drugs and every suppressor tRNA remain unknown.

Five of six scored small molecules have no matched native-stop atlas

Empirical coverage by therapy; dashed rows mean unknown, never zero

absent
CC-90009Clitocine2,6-diaminopurineG418 (geneticin)SJ6986SRI-41315CC-90009: no matched empirical native-stop atlasClitocine: no matched empirical native-stop atlas2,6-diaminopurine: no matched empirical native-stop atlasG418 (geneticin): measured for 1,700 canonical stops in HEK293T cellsSJ6986: no matched empirical native-stop atlasSRI-41315: no matched empirical native-stop atlasNO MATCHED EMPIRICAL ATLASNO MATCHED EMPIRICAL ATLASNO MATCHED EMPIRICAL ATLASMEASURED · HEK293T · 1,700 STOPSNO MATCHED EMPIRICAL ATLASNO MATCHED EMPIRICAL ATLASFILLED = MEASURED NATIVE-STOP BURDENDASHED = UNKNOWN, NOT ZERO

THE SAFETY LAYER IS INCOMPLETE · All suppressor-tRNA designs also lack a qualifying native-stop measurement.

Details & limits
The existing result is downstream ribosome occupancy for G418 in HEK293T cells. It is not protein-extension abundance, cell toxicity or a result in disease-relevant tissue. Each new therapy needs its own treated-versus-control atlas.
3 / armnext measurement

Test one suppressor tRNA against its empty delivery construct: three biological replicates per arm, transcriptome-wide ribosome profiling, no invented pass/fail cutoff.

Direct scope · One suppressor tRNA scaffold in one cell line, measured across that cell line's expressed transcriptome · endpoint · Occupancy past each annotated stop, treated against control, transcriptome-wide